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Scarabée redirige ici Pour les autres significations voir Scarabée homonymie Ne doit pas être confondu avec Scarabaeoide
Scarabaeidae
« Scarabée » redirige ici. Pour les autres significations, voir Scarabée (homonymie).
Ne doit pas être confondu avec Scarabaeoidea.
Les Scarabaeidae, en français Scarabéidés, sont une famille d'insectes coléoptères de la super-famille des Scarabaeoidea. Ils sont connus sous le nom vernaculaire de Scarabée. C'est notamment la famille des dynastes, du scarabée rhinocéros européen, des bousiers et des hannetons.
| Règne | Animalia |
|---|---|
| Sous-règne | Bilateria |
| Infra-règne | Protostomia |
| Super-division | Ecdysozoa |
| Embranchement | Arthropoda |
| Sous-embr. | Hexapoda |
| Classe | Insecta |
| Sous-classe | Pterygota |
| Infra-classe | Neoptera |
| Super-ordre | Holometabola |
| Ordre | Coleoptera |
| Sous-ordre | Polyphaga |
| Infra-ordre | Scarabeiformia |
| Super-famille | Scarabaeoidea |
Famille
Latreille, 1802
Les limites de cette famille sont encore débattues et controversées. Les chercheurs européens en ont une définition plutôt restrictive, alors qu'un certain nombre de chercheurs américains y incorporent bien plus de taxons.
Les antennes des insectes de cette famille sont en massue dont l'extrémité peut s'ouvrir en un éventail de feuillets. L'extrémité de l'abdomen est découverte en raison d'élytres normalement tronqués. La plupart des espèces volent correctement. Plusieurs de ces espèces stridulent en frottant l'extrémité de leurs élytres contre la face dorsale de leur abdomen. Cette vaste famille comprend des coprophages et des saproxylophages nuisibles aux végétaux.
Noms vernaculaires
Le terme de Scarabée provient, via le vieux français « escarbot », du latin scarabaeus, par altération du grec ancien κάραβος / kárabos, « escarbot, scarabée, insecte », également à l'origine du terme carabe. Le terme « scarabée» désigne de façon générale en zoologie l'ensemble de la super-famille des Scarabéoïdes, mais ce terme peut désigner chez certains auteurs, d'une façon plus restrictive, la famille des Scarabaeidae, et même pour certains uniquement les coprophages de cette famille[réf. nécessaire].
Plusieurs groupes, qui peuvent être paraphylétiques, de ces insectes ont des noms vernaculaires tels que les Dynastes ou scarabées rhinocéros, les bousiers, les cétoines, les hannetons... Certaines espèces ont un nom vernaculaire particulier comme le Pique-prune (Osmoderma eremita).
Systématique
La famille a été décrite par l'entomologiste français Pierre André Latreille en 1802.
Liste des sous-familles
Selon le Catalogue of Life (23 juin 2023), cette famille se divise en 28 sous-familles :
- Aclopinae Blanchard, 1850
- Aegialiinae Castelnau, 1840
- Arrow, 1940
- Aphodiinae Leach, 1815
- Janssens, 1946
- Cetoniinae Leach, 1815
- Blanchard, 1845
- † Nikolajev, 1995
- Arrow, 1911
- Dynastinae MacLeay, 1819
- Stebnicka, 1977
- Latreille, 1829
- Burmeister, 1844
- † Nikolajev, 1992
- Melolonthinae MacLeay, 1819
- LeConte, 1861
- Orphninae Erichson, 1847
- Reitter, 1902
- Erichson, 1847
- Howden, 1997
- † Nikolajev, 2000
- Rutelinae MacLeay, 1819
- Scarabaeinae Latreille, 1802
- Sericinae Kirby, 1837
- Erichson, 1847
- Wasmann, 1918
- Trichiinae Kolbe, 1897
- Valginae Schenkling, 1922
Liste des genres
Selon le Catalogue of Life (23 juin 2023), cette famille regroupe 2 590 genres :
- Dejean, 1833
- Blanchard, 1850
- Schmidt, 1909
- Acanthobodilus Dellacasa, 1983
- Janssens, 1937
- Kraatz, 1895
- Prell, 1936
- Britton, 1988
- Erichson, 1848
- Erichson, 1840
- Kraatz, 1888
- Erichson, 1835
- Casey, 1889
- Arrow, 1899
- Ohaus, 1912
- Schmidt, 1914
- Schmidt, 1914
- Acrossus Mulsant, 1842
- Kraatz, 1882
- Westwood, 1841
- Ohaus, 1909
- Bordat & Howden, 1995
- Ratcliffe, 2014
- Schmidt, 1914
- Kraatz, 1883
- Blanchard, 1851
- Adoretus Dejean, 1833
- Brenske, 1893
- Brenske, 1893
- Arrow, 1917
- Blackburn, 1889
- Britton, 1995
- Villiers, 1956
- Latreille, 1807
- Paulian, 1984
- Arrow, 1904
- Westwood, 1845
- Burmeister, 1847
- Dejean, 1833
- Arrow, 1899
- Decelle, 1982
- Burmeister, 1842
- Petrovitz, 1969
- Dellacasa, 1984
- Pittino & Mariani, 1986
- Krikken, 2009
- Bordat, 2019
- Decelle, 1968
- Bordat, 2016
- Bates, 1887
- Reitter, 1895
- Le Peletier & Serville, 1828
- Moser, 1918
- Eschscholtz, 1829
- Brenske, 1896
- Miyake & Iwase, 1991
- Schmidt, 1914
- Mulsant & Rey, 1869
- Dellacasa, Dellacasa & Gordon, 2008
- Agrilinus Mulsant & Rey, 1869
- Báguena, 1930
- (Balthasar, 1963)
- Stebnicka & Howden, 1996
- Král, Mencl & Rakovič, 2021
- Petrovitz, 1980
- Allsopp, 2018
- Lacroix, 2003
- Arrow, 1908
- Deschodt & Scholtz, 2008
- Britton, 1955
- Schenkling, 1921
- Arrow, 1940
- Bordat, 2009
- Petrovitz, 1963
- Janssens, 1940
- Péringuey, 1904
- Allomyrina Arrow, 1911
- Janssens, 1936
- Dupuis & Dechambre, 2001
- Fauvel, 1903
- Burmeister, 1842
- Boucomont, 1923
- Britton, 1978
- Schmidt, 1914
- Péringuey, 1904
- Frey, 1975
- Blanchard, 1850
- Reitter, 1902
- Kolbe, 1895
- Kraatz, 1882
- Blanchard, 1851
- Reitter, 1902
- Dombrow, 2002
- Westwood, 1878
- Kolbe, 1910
- MacLeay, 1819
- Skelley, 2007
- Stebnicka & Skelley, 2003
- Amidorus Mulsant, 1870
- Cambefort, 1981
- Nomura, 1974
- Thomson, 1878
- Bordat, 1999
- Mulsant, 1842
- Fairmaire, 1898
- Huchet, 2000
- Amphimallon Latreille, 1825
- Lansberge, 1874
- Brenske, 1894
- Smith, 2008
- Dechambre & Madge, 1979
- Blanchard, 1842
- Britton, 1990
- Petrovitz, 1969
- Arrow, 1922
- Martinez, 1958
- Antoine, 1989
- Lacroix, 1995
- Kolbe, 1894
- Brême, 1844
- Burmeister, 1842
- Péringuey, 1907
- Curtis, 1835
- Erichson, 1847
- Britton, 1957
- Kolbe, 1892
- Lacroix, 1993
- Blackburn, 1898
- Martinez & Pereira, 1956
- Burmeister, 1844
- Latreille, 1807
- Moser, 1913
- Fischer Von Waldheim, 1824
- Westwood, 1842
- Paulian, 1954
- Howden, 1971
- Burmeister, 1842
- Moser, 1914
- Sharp, 1875
- Ohaus, 1901
- Fairmaire, 1878
- Anomala Samouelle, 1819
- † Fric, 1885
- Moser, 1913
- Arrow, 1908
- Reitter, 1887
- Jameson, Paucar-Cabrera & Solis, 2003
- Paulian, 1952
- Blackwelder, 1944
- Westwood, 1842
- Arrow, 1946
- Mulsant & Rey, 1869
- Fairmaire, 1897
- Saylor, 1940
- Janssens, 1950
- Arrow, 1917
- MacLeay, 1838
- Simonis, 1981
- Rivers, 1889
- Anoplognathus Leach, 1815
- Brullé, 1837
- Arrow, 1939
- Laporte, 1832
- Petrovitz, 1971
- Lacroix, 1993
- Minkina, 2020
- † Gomez Pallerola, 1979
- Medvedev, 1949
- Britton, 1957
- Burmeister, 1842
- Moser, 1918
- Lacroix, 1993
- Eschscholtz, 1818
- Soula, 1998
- † Lu, Ahrens, Bai, Shih & Ren, 2023
- Brenske, 1900
- Burmeister, 1855
- Brême, 1844
- Petrovitz, 1958
- Howden, 1968
- Brenske, 1900
- Britton, 1990
- Kraatz, 1880
- Antoine, 1987
- Harold, 1868
- Péringuey, 1901
- Koshantschikov, 1911
- Schmidt, 1914
- Paulian, 1942
- Aphodius Hellwig, 1798
- Péringuey, 1901
- Endrödi, 1983
- Endrödi, 1964
- Rivers, 1889
- Kolbe, 1905
- LeConte, 1856
- Cartwright, 1952
- Lacroix, 1993
- Blanchard, 1850
- Antoine, 2001
- Kirby, 1818
- Lansberge, 1886
- Brenske, 1903
- Olsoufieff, 1947
- Baraud, 1962
- Schmidt, 1917
- Matthews, 1974
- Montreuil, 2011
- Smith & Evans, 2018
- Gutiérrez, 1952
- Janssens, 1937
- Uliana & Sabatinelli, 2017
- Westwood, 1842
- Morón & Krikken, 1990
- Nikolajev, 1982
- † Minkina, 2020
- Brenske, 1895
- Kolbe, 1905
- Kolbe, 1910
- Brenske, 1897
- Burmeister, 1844
- Paulsen, 2021
- MacLeay, 1819
- Penco & Zubarán, 2013
- Stebnicka & Dellacasa, 2003
- Dombrow, 2003
- Thomson, 1878
- Rossi, 1958
- Péringuey, 1904
- Martinez, 1960
- Stebnicka, 1999
- Brenske, 1894
- Brancsik, 1893
- Makhan, 2006
- Nikolajev, 2000
- (Medvedev, 1949)
- Petrovitz, 1961
- Bates, 1888
- Rojkoff, 2013
- Aspilus Westwood, 1874
- Erichson, 1847
- Martinez, 1957
- Brenske, 1898
- Westwood, 1874
- Westwood, 1843
- Britton, 1990
- Péringuey, 1907
- Schürhoff, 1937
- Petrovitz, 1973
- Ataenius Harold, 1867
- Petrovitz, 1954
- Bedel, 1892
- † Meunier, 1898
- Ateuchus Weber, 1801
- Lacroix, 1998
- Erichson, 1835
- González-Alvarado, Molano-Rendón & Vaz-de-Mello, 2019
- Kraatz, 1898
- Distant, 1911
- Philips & Bell, 2008
- Strand, 1942
- Burmeister, 1847
- Burmeister, 1841
- Guérin-Méneville, 1838
- White, 1859
- Decelle, 1973
- Brenske, 1900
- Klug, 1838
- Hutchinson & Moeseneder, 2019
- Deschodt & Davis, 2018
- Mencl, 2008
- Petrovitz, 1958
- (Balthasar, 1942)
- Howden & Storey, 1992
- Britton, 1978
- Brenske, 1897
- Krikken, 2018
- Mikšić, 1976
- Thomson, 1880
- Machatschke, 1970
- Thomson, 1877
- † Nikolajev, 2005
- Paucar-cabrera, 2003
- Fairmaire, 1898
- Gordon & Skelley, 2007
- Paulian, 1986
- Thomson, 1880
- Allsopp, 2022
- Ratcliffe, 1981
- Blanchard, 1850
- Endrödi, 1956
- Schein, 1957
- Chalumeau, 1983
- Pereira & Vulcano & Martinez, 1960
- Harold, 1869
- Dombrow, 2005
- Péringuey, 1904
- Vaz-de-Mello, 2008
- Fairmaire, 1898
- Fairmaire, 1893
- Machatschke, 1970
- White, 1841
- Mulsant & Rey, 1869
- Antoine, 2003
- Lacroix, 1993
- Paulian, 1961
- Péringuey, 1902
- Fairmaire, 1898
- Schmidt, 1914
- Burmeister, 1842
- Lacroix, 1998
- Lacroix, 1994
- Blanchard, 1850
- Péringuey, 1902
- Reitter, 1903
- Endrödi, 1973
- Dellacasa & Dellacasa, 2005
- Adam, 1994
- Mulsant & Rey, 1869
- Paulian, 1975
- Fisher Von Waldheim, 1829
- Harold, 1868
- Matthews, 1974
- Westwood, 1848
- Ruter, 1978
- Pittino & Mariani, 1986
- Halffter, 1958
- Şenyüz, 2017
- Burmeister, 1842
- Hope, 1837
- Balthasar, 1932
- Kraatz, 1890
- Koshantschikov, 1913
- Kolbe, 1894
- Brenske, 1896
- Kraatz, 1896
- Thomson, 1858
- Brenske, 1898
- Waterhouse, 1881
- Guérin-Méneville, 1831
- Vaz-de-Mello, 2008
- Blanchard, 1851
- Baraud, 1986
- Berg, 1898
- Ricchiardi, 2017
- Burmeister, 1842
- Landin, 1960
- Özdikmen & Demir, 2008
- Martinez, 1952
- Howden & Storey, 1992
- Dejan, 1833
- Sabatinelli & Pontuale, 1998
- Nomura, 1970
- Berg, 1898
- Dombrow, 2003
- Mikšić, 1955
- Harold, 1869
- Harold, 1869
- Blackburn, 1892
- Brenske, 1898
- Burmeister, 1844
- Balthasar, 1964
- Caccobius Thomson, 1859
- Paulian, 1984
- Lewis, 1895
- Hope, 1841
- Lacroix, 1998
- Motschulsky, 1859
- † Krell, 2011
- Calicnemis Laporte, 1832
- Gordon & Skelley, 2007
- Blackburn, 1895
- Lacroix, 1998
- Burmeister, 1842
- Blanchard, 1851
- Péringuey, 1901
- Blanchard, 1851
- Ohaus, 1903
- White, 1845
- Moser, 1916
- Brenske, 1894
- Mohnike, 1873
- Dellacasa, 1984
- Blanchard, 1850
- Bates, 1888
- Bourgoin, 1917
- Blanchard, 1850
- Howden, 1970
- Ricchiardi, 2000
- Paulian, 1986
- Branco, 1989
- Erichson, 1847
- Brenske, 1900
- Lacroix, 2005
- Lacroix, 2002
- Hope, 1831
- Kirby, 1827
- Decelle, 1979
- Brenske, 1900
- Stebnicka & Howden, 1995
- Saylor, 1938
- Erichson, 1847
- Chapin, 1934
- Davis, Scholtz & Harrison, 1999
- Canthon Hoffmannsegg, 1817
- Chapin, 1930
- Paulian, 1939
- MacLeay, 1871
- Paulian, 1939
- Bordat, 2022
- Lacroix, 1994
- Lacroix, 1999
- Tesař, 1945
- Bezděk, 2016
- Endrödi, 1974
- Mikšić, 1976
- Özdikmen, 2009
- Özdikmen, 2009
- Islas, 1958
- Ohaus, 1909
- Lacroix, 1999
- Balthasar, 1967
- Hope, 1837
- Solier, 1851
- Coca-Abia & Robbins, 2006
- Sharp, 1875
- Ratcliffe & Cave, 2010
- † Woolley, 2016
- Burmeister, 1842
- Fairmaire, 1884
- Krikken, 1976
- Guérin-Méneville, 1840
- Dellacasa & Gordon & Dellacasa, 1998
- Westwood, 1842
- Brenske, 1898
- Moser, 1920
- Baraud, 1987
- Ceraspis Le Peletier & Serville, 1828
- Dombrow, 2002
- Kolbe, 1899
- Arrow, 1948
- Ohaus, 1911
- Arrow, 1908
- Quedenfeldt, 1884
- Gestro, 1893
- Koçak & Kemal, 2008
- Ohaus, 1912
- Cetonia Fabricius, 1775
- Moser, 1917
- Heller, 1921
- Schaum, 1862
- Westwood, 1847
- Medvedev, 1949
- Moser, 1914
- Arrow, 1937
- Burmeister, 1844
- Ohaus, 1905
- Burmeister, 1846
- Heller, 1903
- Dejean, 1833
- Heller, 1902
- Burmeister, 1844
- Chalcosoma Hope, 1837
- Burmeister, 1842
- Mikšić, 1970
- Casey, 1915
- Burmeister, 1842
- Brenske, 1898
- Moser, 1919
- Blanchard, 1850
- Brenske, 1899
- Kolbe, 1904
- Dejean, 1821
- Le Peletier & Serville, 1828
- Saylor, 1937
- Britton, 1990
- MacLeay, 1864
- Britton, 1957
- Westwood, 1842
- Lansberge, 1875
- Hope, 1837
- Britton, 1957
- Hope, 1840
- Vardal & Forshage, 2010
- Morón & Nogueira, 1998
- Lacroix, 1993
- Croissandeau, 1893
- Legrand, 2004
- Smith & Mondaca, 2016
- Sharp, 1877
- Burmeister, 1842
- Moser, 1924
- Chilothorax Motschulsky, 1859
- Reitter, 1905
- Kraatz, 1891
- Soula, 2008
- Bates, 1889
- MacLeay, 1819
- Tarasov, 2008
- Blanchard, 1850
- Fairmaire, 1905
- Bates, 1889
- Kraatz, 1880
- Kirby, 1828
- Burmeister, 1844
- Burmeister, 1844
- Kraatz, 1880
- Kraatz, 1880
- Westwood, 1842
- Kolbe, 1897
- Kirby, 1828
- Le Peletier & Serville, 1828
- Brenske, 1897
- Burmeister, 1847
- Soula, 2005
- Schmidt, 1914
- Latreille, 1829
- Schein, 1956
- Burmeister, 1840
- Laporte, 1832
- Cambefort, 1984
- Burmeister, 1842
- Lea, 1914
- Burmeister, 1842
- Motschulsky, 1857
- Schoch, 1898
- Ohaus, 1941
- Burmeister, 1842
- Erichson, 1847
- Lacroix, 1999
- Fairmaire, 1904
- Krikken, 2009
- Li & Yang, 1999
- Frey, 1970
- Arrow, 1908
- Semenov, 1903
- Schmidt, 1914
- Kirby, 1827
- Kolbe, 1910
- Motschulsky, 1868
- Decelle, 1979
- Paulian, 1991
- Lacroix, 1996
- Kolbe, 1894
- Burgeon, 1946
- Péringuey, 1902
- Kolbe, 1895
- Burmeister & Schaum, 1841
- Moser, 1913
- Hope, 1837
- Schmidt, 1914
- Schaum, 1841
- Horn, 1876
- Prell, 1933
- Hope, 1831
- Ohaus, 1910
- Ratcliffe & Hardy, 2005
- Clément, 1958
- Paulian, 1942
- Mulsant, 1842
- Blanchard, 1850
- Britton, 1987
- Erichson, 1843
- Blackburn, 1898
- Blanchard, 1850
- Brenske, 1897
- Lacroix, 2008
- Lacroix, 1993
- Strand, 1928
- Lacroix, 1997
- White, 1845
- Fuhrmann, 2012
- Kolbe, 1884
- Fauvel, 1903
- Erichson, 1847
- Brenske, 1900
- Kolbe, 1892
- Boucomont, 1920
- Mulsant, 1842
- Copris Geoffroy, 1762
- Reiche, 1841
- Olsoufieff, 1924
- Balthasar, 1938
- Harold, 1859
- Burmeister, 1846
- Burmeister, 1847
- Coptomia Burmeister, 1842
- Pouillaude, 1919
- Hope, 1833
- Hope, 1845
- Moser, 1918
- Kolbe, 1892
- Klug, 1845
- Prell, 1936
- Kraatz, 1880
- Saunders, 1852
- Kraatz, 1898
- Kolbe, 1897
- Given, 1952
- Schürhoff, 1933
- Cotalpa Burmeister, 1844
- Burmeister, 1842
- Kolbe, 1895
- Schmidt, 1914
- Blanchard, 1851
- Knoch, 1801
- Brenske, 1903
- † Nikolajev, 2004
- † Nikolajev, 1993
- † Nikolajev, 1998
- † Nikolajev, 2007
- † Nikolajev, 1995
- † Nikolajev, 1995
- † Nikolajev, 1998
- Bordat, 2009
- Balthasar, 1942
- Burmeister, 1847
- MacLeay, 1819
- Gordon & Skelley, 2007
- Kraatz, 1888
- Britton, 1992
- Lea, 1916
- Krikken, 1976
- Westwood, 1874
- MacLeay, 1838
- Cyclocephala Dejean, 1821
- Klug, 1855
- Burmeister, 1844
- Schein, 1958
- Ohaus, 1923
- Kirby, 1827
- Sharp, 1875
- Waterhouse, 1867
- Burmeister, 1847
- Westwood, 1842
- Fischer Von Waldheim, 1823
- Brenske, 1900
- Lesne, 1900
- Burmeister, 1842
- Reitter, 1903
- Brenske, 1897
- Fairmaire, 1895
- Kolbe, 1895
- Brenske, 1900
- Arrow, 1917
- Howden & Storey, 2000
- Prokofiev, 2013
- Dechambre, 2001
- Bordat & Howden, 1995
- MacLeay, 1819
- Moser, 1913
- Reitter, 1902
- Le Peletier & Serville, 1828
- Endrödi, 1952
- Kolbe, 1904
- Allsopp, 2022
- Lacroix, 1993
- Lacroix, 2005
- Lacroix, 1999
- Sabatinelli & Pontuale, 1998
- Vaz-de-Mello, 2008
- Gordon & Skelley, 2007
- Erichson, 1848
- Fairmaire, 1900
- Pereira, 1949
- Eschscholtz, 1822
- Harold, 1867
- Balthasar, 1961
- Burmeister, 1847
- Perty, 1830
- Dechambre, 2006
- Lacroix, 1997
- Schein, 1959
- Dombrow, 2002
- Arrow, 1941
- Moser, 1915
- Ruter, 1964
- Ocampo & Mondaca, 2012
- Alexis & Delpont, 1999
- Thomson, 1878
- Arrow, 1907
- Gistel, 1857
- Brenske, 1896
- De Wailly, 1950
- Parry, 1849
- Brown, 1929
- Harold, 1869
- Gordon & Skelley, 2007
- Newman, 1840
- Péringuey, 1904
- Erichson, 1847
- Burmeister, 1844
- Horn, 1887
- Lansberge, 1886
- Mulsant, 1842
- Gestro, 1888
- Waterhouse, 1905
- Burmeister, 1844
- Brenske, 1895
- Blanchard, 1850
- Burmeister, 1855
- Harris, 1826
- Moser, 1924
- Le Peletier & Serville, 1828
- Gory & Percheron, 1833
- Burmeister, 1847
- Dichotomius Hope, 1838
- Kraatz, 1885
- Le Peletier & Serville, 1828
- Frolov & Scholtz, 2003
- Burmeister, 1844
- Reitter, 1903
- Dicronocephalus Hope, 1831
- Hope, 1837
- Orbigny, 1896
- Wood-mason, 1878
- Thomson, 1877
- Balthasar, 1959
- Gutiérrez, 1949
- Britton, 1990
- Lacroix, 1998
- Reiche, 1859
- Dilochrosis Thomson, 1878
- Bates, 1888
- Bordat, 2009
- Casey, 1889
- Lacroix, 1989
- Lacroix, 1989
- Felsche, 1906
- Matthews, 1974
- Hope, 1842
- Gerstaecker, 1882
- Le Peletier & Serville, 1828
- Fairmaire, 1878
- Thomson, 1858
- Hope, 1843
- Kolbe, 1892
- Gory & Percheron, 1833
- Kirby, 1837
- Boheman, 1857
- Burmeister, 1842
- Burmeister, 1842
- Burmeister, 1842
- Harold, 1873
- Lacroix, 1993
- Lacroix, 1993
- Balthasar, 1941
- Péringuey, 1904
- Péringuey, 1904
- Dombrow, 2003
- Lacroix, 1998
- Kolbe, 1892
- Branco, 1989
- Burmeister, 1842
- Blanchard, 1845
- Brenske, 1897
- Schmidt, 1914
- Péringuey, 1901
- Kirby, 1828
- Boucomont, 1921
- Deschodt, Davis & Scholtz, 2016
- Bordat, 2009
- Carne, 1958
- Deschodt & Scholtz, 2008
- Burmeister, 1842
- Dynastes MacLeay, 1819
- Harold, 1869
- Blackburn, 1898
- Dechambre, 1979
- Kraatz, 1880
- Erichson, 1847
- Redtenbacher, 1868
- Ohaus, 1915
- Neita & Ocampo, 2012
- Burmeister, 1842
- Gene, 1836
- Moser, 1919
- Franz, 1955
- Péringuey, 1907
- Schmidt, 1914
- Lacroix, 1998
- Brenske, 1895
- Erichson, 1847
- De Wailly, 1950
- Erichson, 1847
- Gerstaecker, 1871
- Sharp, 1877
- Erichson, 1847
- Endrödi, 1971
- Dejean, 1833
- Lansberge, 1880
- Kolbe, 1892
- Chalumeau, 1981
- Scholtz & Howden, 1987
- Hope, 1837
- Dalle Torre, 1912
- Lacroix, 2002
- Britton, 1990
- Lansberge, 1874
- Lucas, 1859
- Dechambre, 1999
- Erichson, 1847
- Moser, 1913
- Kolbe, 1894
- Eocorythoderus Marumaya, 2012
- Barbero, Palestrini & Roggero, 2009
- Arrow, 1908
- Carne, 1956
- Medvedev, 1951
- Olsoufieff, 1947
- Reitter, 1903
- Blanchard, 1851
- Dejean, 1833
- Bates, 1904
- White, 1841
- Roggero, Barbero & Palestrini, 2015
- Reiche, 1841
- Moser, 1917
- Dejean, 1833
- Carne, 1957
- Fairmaire, 1880
- Tagawa, 1941
- Burmeister, 1842
- Hutchinson & Allsopp, 2021
- Lacroix, 1994
- Mulsant, 1851
- Ohaus, 1910
- Ohaus, 1910
- Kolbe, 1914
- Kolbe, 1914
- Burmeister, 1844
- Lacroix, 2006
- Burmeister & Schaum, 1840
- Burmeister, 1847
- Brenske, 1899
- Burmeister, 1842
- Preiss, 1902
- Schmidt, 1917
- Mulsant & Rey, 1870
- Zídek & Pokorný, 2011
- Mulsant & Rey, 1869
- Péringuey, 1904
- Péringuey, 1904
- Burmeister, 1842
- Burmeister & Schaum, 1840
- Arrow, 1907
- Burmeister, 1842
- Prell, 1912
- Brullé, 1837
- Waterhouse, 1875
- White, 1839
- Burmeister, 1840
- Mulsant & Rey, 1869
- Bates, 1888
- Bordat, 2003
- Dellacasa, 1983
- Lacroix, 1997
- Fairmaire, 1899
- Kolbe, 1894
- Burgeon, 1946
- Casey, 1915
- Mikšić, 1976
- Ohaus, 1909
- Euoniticellus Janssens, 1953
- Balthasar, 1959
- Dellacasa, 1983
- Dellacasa, Gordon & Dellacasa, 2016
- Le Peletier & Serville, 1828
- Brown, 1927
- Hinton, 1936
- Stebnicka & Howden, 1996
- Eupatorus Burmeister, 1847
- Duvivier, 1892
- Brenske, 1897
- Burmeister, 1842
- Mulsant, 1842
- Burmeister, 1842
- Semenov, 1890
- Westwood, 1867
- Marseul, 1867
- Blanchard, 1851
- Blanchard, 1850
- Burmeister, 1842
- Lacroix, 2006
- Brenske, 1898
- Dalman, 1824
- Moser, 1908
- Branco, 1989
- Thomson, 1880
- Reitter, 1896
- Waterhouse, 1880
- Waterhouse, 1882
- Martinez, 1967
- Semenov, 1890
- Castelnau, 1840
- Szito, 1993
- Soula, 1998
- Endrödi, 1960
- Keith & Reichenbach, 2018
- Soula, 2002
- Carne, 1958
- Reitter, 1902
- Reitter, 1903
- Motschulsky, 1859
- Soula, 2002
- Britton, 1979
- Soula, 2002
- Bordat, Mencl & Rakovič, 2012
- Smith & Mondaca, 2015
- Martinez, 1982
- Lacroix, 1997
- Paulian, 1987
- Keith & Montreuil, 2004
- Blanchard, 1850
- Vaz-de-Mello, 2008
- Dellacasa, Dellacasa & Gordon, 2007
- Coca-Abia, 2003
- Gordon & Skelley, 2007
- Stebnicka, 1999
- Bertoloni, 1852
- Howden, 1961
- Balthasar, 1938
- Blackburn, 1892
- Kolbe, 1894
- Bordat, 2022
- Burmeister, 1842
- Antoine, 2005
- Péringuey, 1904
- Ratcliffe, Jameson & Zorn, 2018
- Péringuey, 1907
- Janssens, 1940
- Nomura, 1973
- Brenske, 1897
- Kirby, 1818
- Costa Lima, 1940
- Vaz-de-Mello, 2008
- Westwood, 1874
- Kirby, 1825
- Gordon & Skelley, 2007
- Gordon & Pittino, 1992
- Geotrogus Guérin-Méneville, 1842
- Endrödi, 1974
- Howden, 1988
- Howden, 1980
- Janssens, 1937
- Balthasar, 1933
- Matsumoto, 2014
- Moser, 1924
- Brenske, 1899
- Lin, 1980
- Lacroix, 1993
- Frey, 1974
- Blackburn, 1898
- Schoch, 1896
- Burmeister, 1842
- Brenske, 1899
- Westwood, 1838
- Brenske, 1895
- Bates, 1889
- Mikšić, 1976
- Reiche, 1860
- Reiche, 1856
- Kirby, 1825
- Arrow, 1914
- Erichson, 1847
- Casey, 1915
- Lansberge, 1887
- Kraatz, 1880
- Krajčik, 2009
- Gnorimus Le Peletier & Serville, 1828
- Endrödi, 1982
- Janson, 1881
- Goliathus Lamarck, 1801
- Thomson, 1878
- Golofa Hope, 1837
- Schmidt, 1914
- Dellacasa, Dellacasa & Gordon, 2012
- Dellacasa, Dellacasa & Gordon, 2012
- Reitter, 1892
- Stebnicka & Howden, 1996
- Burmeister, 1844
- Kolbe, 1910
- Arrow, 1911
- Quedenfeldt, 1888
- Ratcliffe, 2010
- Skelley, Dellacasa & Dellacasa, 2009
- Péringuey, 1902
- Ocampo & Mondaca, 2012
- Lacroix, 1997
- Kolbe, 1895
- Hutchinson & Moeseneder, 2013
- Ziani, 2002
- Rakovič, 1981
- Prokofiev, 2016
- Mlambo, Scholtz & Deschodt, 2019
- Lacroix, 1993
- Brullé, 1837
- Schaeffer, 1927
- Péringuey, 1904
- Masumoto & Kiuchi, 2001
- Neervoort Van De Poll, 1886
- Ratcliffe, 2019
- Casey, 1915
- Gymnetis MacLeay, 1819
- Blanchard, 1851
- Dejean, 1833
- Arrow, 1910
- Gymnopleurus Illiger, 1803
- Linell, 1896
- Moser, 1914
- Brenske, 1896
- Lansberge, 1874
- Brenske, 1893
- Kraatz, 1898
- Britton, 1990
- Britton, 1987
- Howden, 1995
- Kraatz, 1892
- Morón & Nogueira, 2007
- Frey, 1967
- Cambefort, 1978
- Roggero, Dierkens, Barbero & Palestrini, 2017
- Halffter & Martinez, 1977
- Hope, 1837
- Kolbe, 1910
- Arrow, 1908
- Semenov, 1890
- Péringuey, 1901
- Harold, 1861
- Bordat, 2003
- Chalumeau, 1981
- Gordon & Skelley, 2007
- Boucomont, 1914
- Britton, 1957
- Harpina Burmeister, 1844
- Burmeister, 1847
- Sharp, 1877
- Kolbe, 1894
- Thomson, 1881
- Orbigny, 1915
- Heliocopris Hope, 1837
- Baraud & Nicolas, 1966
- Krikken, 1982
- Brenske, 1897
- Baraud, 1986
- Mikšić, 1970
- Kraatz, 1880
- Tangelder & Krikken, 1982
- Reitter, 1897
- Schmidt, 1914
- Reiche, 1860
- Mikšić, 1974
- Kraatz, 1880
- Mikšić, 1972
- Burmeister, 1842
- Kolbe, 1910
- Mikšić, 1963
- Mulsant, 1842
- Brenske, 1898
- Motschulsky, 1857
- Seilliere, 1910
- Antoine, 1989
- Burmeister, 1844
- Soula, 2002
- Burmeister, 1842
- Péringuey, 1901
- Albers, 1852
- Kolbe, 1900
- Martinez, 1948
- Bourgoin, 1919
- Ohaus, 1909
- Moser, 1911
- Dupuis & Dechambre, 2008
- Burmeister, 1847
- Kolbe, 1900
- Gillet, 1911
- Heteronychus Dejean, 1833
- Guérin-Méneville, 1830
- Burmeister, 1842
- Blanchard, 1851
- Valck Lucassen, 1933
- Westwood, 1842
- Moser, 1917
- Brenske, 1899
- Schaum, 1844
- Dupont, 1832
- Dupuis, 2006
- Antoine, 2002
- Moser, 1918
- Krikken, 1978
- Fairmaire, 1891
- Olivier, 1789
- Fairmaire, 1901
- Endrödi, 1978
- Burmeister, 1855
- Fairmaire, 1886
- Sabatinelli, 1983
- Howden & Endrodi, 1978
- † Nikritin, 1977
- Péringuey, 1904
- Martinez & Pereira, 1956
- Hope, 1838
- Reitter, 1889
- Brenske, 1894
- Martinez, 1949
- Brenske, 1891
- Brenske, 1896
- Britton, 1978
- Hope, 1837
- Blanchard, 1851
- Brenske, 1897
- Janssens, 1932
- Soula, 2006
- Burmeister, 1842
- Burmeister, 1847
- Kolbe, 1894
- Ohaus, 1901
- MacLeay, 1871
- Guérin-Méneville, 1839
- Kolbe, 1910
- Lacroix, 1998
- Ohaus, 1898
- Kolbe, 1895
- Brenske, 1899
- Künckel D'Herculais, 1887
- Hoplia Illiger, 1803
- Blanchard, 1850
- Kolbe, 1904
- Blanchard, 1851
- Harold, 1869
- Fairmaire, 1904
- Nomura, 1974
- Bates, 1904
- Laporte, 1832
- Thomson, 1880
- Schürhoff, 1933
- Moser, 1918
- Prell, 1933
- Burgeon, 1946
- Stebnicka, 2000
- Dellacasa, Dellacasa & Gordon, 2015
- Endrödi, 1974
- Fairmaire, 1897
- Arrow, 1911
- Hardy, 1978
- Arnaud, 1993
- Lacroix, 1993
- Palestrini & Giacone, 1988
- Quedenfeldt, 1884
- Dejean, 1833
- Brenske, 1898
- Péringuey, 1901
- Péringuey, 1904
- † Nikolajev, 1996
- Kolbe, 1904
- Arrow, 1899
- Ohaus, 1910
- Reitter, 1890
- Eschscholz, 1830
- Arrow, 1899
- Fairmaire, 1878
- Kraatz, 1880
- Blackburn, 1895
- Balthasar, 1938
- Britton, 1978
- Brenske, 1897
- Howden, 1968
- LeConte, 1861
- Burmeister, 1840
- Martinez, 1953
- Rössner, 2018
- Gory & Percheron, 1833
- Britton, 1986
- Péringuey, 1904
- Péringuey, 1904
- Britton, 1987
- Arrow, 1916
- Arrow, 1916
- Arrow, 1932
- Arrow, 1910
- Péringuey, 1902
- Heller, 1916
- Stebnicka, 1997
- Montreuil & Lacroix, 2020
- Péringuey, 1902
- Le Peletier & Serville, 1828
- Thomson, 1857
- Dechambre, 1979
- Mencl, Král & Rakovič, 2019
- Krikken, 2009
- Krikken, 2009
- Ricchiardi, 2012
- Endrödi, 1976
- Fairmaire, 1893
- Smith & Mondaca, 2015
- Nikolajev, 2006
- Ratcliffe, 2003
- Delgado & Morón, 1991
- Gordon & Skelley, 2007
- Pacheco & Vaz-de-Mello, 2019
- Gestro, 1874
- Kraatz, 1892
- Burmeister, 1842
- Krikken, 1978
- Kraatz, 1880
- Kolbe, 1897
- Pereira & Martinez, 1960
- Mannerheim, 1828
- Burmeister, 1855
- Fairmaire, 1889
- Roth, 1851
- Thomson, 1880
- Balthasar, 1967
- Dellacasa, Gordon & Dellacasa, 2003
- Schürhoff, 1937
- Paulian, 1976
- Cambefort, 1975
- Dierkens, Moretto & Génier, 2017
- Lacroix, 1993
- Saunders, 1839
- Dalla Torre, 1913
- † Nikolajev, 2005
- Lacroix, 2005
- Frey, 1965
- Péringuey, 1902
- Minkina, 2017
- Perissinotto, 2020
- Lacroix, 2011
- Lacroix, 2009
- Delpont, 1996
- Janssens, 1940
- Péringuey, 1902
- Nagai, 1984
- Allsopp, 2022
- Péringuey, 1902
- Jacobson, 1906
- Lacroix, 1993
- Kim, 1996
- Péringuey, 1902
- Schmidt, 1914
- Péringuey, 1904
- Branco, 1991
- Nikolajev & Kabakov, 1977
- Péringuey, 1902
- Lacroix, 1989
- Roggero, Barbero & Palestrini, 2017
- Mulsant & Rey, 1869
- Sharp, 1873
- Lacroix, 1997
- Erichson, 1847
- Reitter, 1889
- Keith, 2000
- Keith, 2007
- Hoffmannsegg, 1817
- Krikken, 1980
- Brenske, 1897
- Ritsema, 1888
- Keith, 2007
- Balthasar, 1963
- Semenov-Tian-Shanskii & Medvedev, 1936
- Blanchard, 1851
- Erichson, 1847
- Brenske, 1897
- Reitter, 1899
- Reitter, 1913
- Moser, 1921
- Krikken, 2009
- Lacroix, 1993
- Kolbe, 1907
- Kolbe, 1894
- Gordon & Skelley, 2007
- Rakovič, 1981
- MacLeay, 1863
- Britton, 1986
- Fabrizi, Eberle & Ahrens, 2019
- Vaz-de-Mello, 2008
- Balthasar, 1966
- Lepidiota Kirby, 1828
- Kolbe, 1894
- Nikolajev, 1979
- Kolbe, 1894
- Moser, 1914
- Le Peletier & Serville, 1828
- Le Peletier & Serville, 1828
- Schein, 1959
- Moser, 1914
- Baraud & Branco, 1991
- Dombrow, 2001
- Saylor, 1935
- Howden, 2003
- Péringuey, 1904
- Burmeister, 1842
- Dejean, 1833
- Brenske, 1892
- MacLeay, 1819
- Erichson, 1847
- Reitter, 1892
- Erichson, 1835
- Gutiérrez, 1946
- Martinez, 1953
- Prell, 1933
- Burmeister, 1847
- Mulsant & Rey, 1869
- Lacroix, 1999
- Rakovič & Mencl, 2012
- Petrovitz, 1961
- Guérin-Méneville, 1830
- Pittino, 2006
- Hutchinson & Allsopp, 2022
- Burmeister, 1842
- Motschulsky, 1859
- Kolbe, 1892
- Guérin-Méneville, 1830
- Arrow, 1917
- Schaum, 1844
- Bates, 1889
- Blanchard, 1851
- † Nikolajev, 1992
- † Nikolajev, 1992
- Waterhouse, 1891
- † Montreuil, Génier & Nel, 2010
- Ohaus, 1917
- Schmidt, 1914
- Kolbe, 1914
- Kolbe, 1897
- Martinez, 1951
- Stebnicka, 1999
- Gory & Percheron, 1833
- Kolbe, 1895
- Burmeister, 1847
- Endrödi, 1980
- Frey, 1974
- Janssens, 1938
- Westwood, 1842
- Perissinotto, Clennell & Beinhundner, 2019
- Reitter, 1892
- Mulsant & Rey, 1869
- Péringuey, 1901
- Frey, 1969
- Mikšić, 1968
- Olsoufieff, 1940
- Martinez, 1972
- Balthasar, 1964
- Westwood, 1875
- Özdikmen, 2009
- Gordon & Skelley, 2007
- Brême, 1844
- Kraatz, 1880
- Lacroix, 1993
- Britton, 1990
- MacLeay, 1819
- Schürhoff, 1935
- Ohaus, 1910
- Macrodactylus Latreille, 1825
- Westwood, 1842
- Westwood, 1874
- Hoffmannsegg, 1817
- Krikken, 1977
- Brenske, 1898
- Lacroix, 1997
- Morón, 1983
- Guérin-Méneville, 1844
- Ohaus, 1905
- Bordat, Dellacasa & Dellacasa, 2000
- Péringuey, 1908
- Strand, 1934
- Gordon & Skelley, 2007
- Lacroix, 1998
- Paulian, 1992
- Ratcliffe & Romé, 2019
- Chalumeau & Gruner, 1976
- Lea, 1919
- MacLeay, 1819
- Montreuil & Keith, 2022
- Lacroix, 1993
- Péringuey, 1908
- Lacroix, 1993
- Mulsant, 1871
- Martinez, 1961
- Heller, 1891
- Arrow, 1941
- Lacroix, 2013
- Blanchard, 1850
- Paulian, 1974
- Moser, 1919
- Moser, 1919
- Castelnau, 1840
- Moser, 1924
- Kirby, 1827
- Thomson, 1880
- Coquerel, 1866
- Chalumeau & Özdikmen, 2006
- Vaz-de-Mello, 2008
- Arrow, 1919
- Hirasawa, 1992
- Gunter & Weir, 2017
- Dombrow, 2002
- Lansberge, 1882
- Alexis & Delpont, 2001
- Lacroix, 1993
- Morón & Nogueira, 1998
- Antoine, 1997
- Bates, 1904
- Mulsant, 1870
- Hope, 1837
- Keith, 2001
- Hope, 1837
- Dechambre, 1976
- Britton, 1987
- Janssens, 1937
- (Reitter, 1903)
- Schürhoff, 1933
- Britton, 1978
- Megasoma Kirby, 1825
- Kakizoe, Maruyama & Masumoto, 2019
- Landin, 1974
- Reitter, 1892
- Waterhouse, 1891
- Eschscholtz, 1822
- Balthasar, 1939
- Burmeister, 1855
- Lacroix, 1997
- Kraatz, 1900
- Fairmaire, 1881
- Brenske, 1899
- Melanocanthon Halffter, 1958
- Lin, 1980
- Melinopterus Mulsant, 1842
- Wollaston, 1869
- Melolontha Fabricius, 1775
- † Heer, 1847
- De Wailly, 1950
- Reitter, 1901
- Harold, 1868
- Arrow, 1941
- Castelnau, 1840
- Brenske, 1897
- Krikken, 1982
- Lacroix, 1993
- Gordon & Skelley, 2007
- Thomson, 1880
- Ohaus, 1905
- Motschulsky, 1863
- Brenske, 1897
- Morón, 1987
- Kolbe, 1914
- Rakovič & Král, 1997
- † Ponomarenko, 1977
- Waterhouse, 1878
- Paulian, 1939
- Machatschke, 1957
- Kraatz, 1880
- Valck Lucassen, 1933
- Arrow, 1911
- Bates, 1889
- Kolbe, 1910
- Lacroix, 2008
- Westwood, 1842
- Britton, 1978
- Frolov & Akhmetova, 2017
- Ruter, 1969
- Lacroix, 1997
- Brown, 1931
- Arrow, 1911
- Pouillaude, 1917
- Moser, 1910
- Blanchard, 1851
- Britton, 1957
- Balthasar, 1958
- Jákl, 2009
- Burmeister, 1844
- Kraatz, 1885
- Petrovitz, 1967
- Gutiérrez, 1951
- Blanchard, 1842
- Baraud, 1991
- Kolbe, 1910
- Kraatz, 1890
- Péringuey, 1902
- Burmeister, 1844
- Kraatz, 1880
- Lea, 1920
- Bates, 1904
- Arrow, 1908
- Brenske, 1894
- Nikolajev, 1979
- Pittino, 2006
- Landin, 1974
- Burmeister, 1855
- Brenske, 1900
- Kraatz, 1883
- Ohaus, 1911
- Montreuil & Keith, 2022
- Péringuey, 1901
- Arrow, 1901
- Nikolajev, 2007
- Kirby, 1823
- Casey, 1915
- Martinez, 1964
- Balthasar, 1963
- Arrow, 1944
- Soula, 1998
- Arrow, 1902
- Lacroix, 1998
- † Wickham, 1914
- Lacordaire, 1855
- Ruter, 1953
- Reitter, 1902
- Brenske, 1903
- Thomson, 1859
- Burmeister, 1844
- Ritsema, 1888
- MacLeay, 1821
- Fairmaire & Germain, 1860
- Kraatz, 1897
- Antoine, 1989
- † Nikolajev, 2004
- Le Peletier & Serville, 1828
- Lansberge, 1874
- Erichson, 1847
- Howden & Storey, 2000
- Gunter & Weir, 2017
- Roggero, Dierkens, Barbero & Palestrini, 2017
- Kraatz, 1880
- Janson, 1905
- Ramírez-Ponce, 2015
- Grossi & Vaz-de-Mello, 2015
- Ahrens, Lukic & Liu, 2023
- Ruter, 1965
- Semenov & Medvedev, 1927
- Huang & Chen, 2016
- Nomura & Nakane, 1951
- Soula, 2002
- Lacroix, 1993
- Vauloger, 1898
- Broun, 1904
- Neervoort Van De Poll, 1886
- Berg, 1881
- Martinez, 1975
- Ruter, 1964
- Bourgoin, 1920
- Burmeister, 1840
- Balthasar, 1955
- Martinez, 1952
- Wasmann, 1900
- Petrovitz, 1962
- Burmeister, 1842
- Perissinotto, 2020
- Hirasawa, 1992
- Scholtz & Howden, 1987
- Deschodt & Davis, 2017
- Péringuey, 1902
- Burmeister, 1844
- Kolbe, 1894
- Britton, 1978
- Westwood, 1842
- Howden, 2003
- Stebnicka, 1999
- Dupont, 1835
- Moeseneder & Hutchinson, 2016
- Morón & Nogueira, 1998
- Koshantschikov, 1912
- Deschodt & Scholtz, 2008
- Brenske, 1900
- Saylor, 1938
- Arrow, 1906
- Arrow, 1917
- Bordat, 1990
- Burgeon, 1945
- Perissinotto, 2017
- Neervoort Van De Poll, 1886
- Lacroix, 1997
- Sharp, 1878
- Landin, 1974
- Ratcliffe & Mico, 2001
- Carne, 1985
- Carne, 1957
- Dellacasa, 1984
- Clément, 1986
- Martinez, 1953
- Paulian & Villiers, 1941
- Blackburn, 1890
- Heller, 1896
- Baraud, 1965
- Yamaya, 2010
- Minkina, 2023
- Alexis & Delpont, 1998
- Carne, 1957
- Péringuey, 1901
- Lacroix, 2015
- Lucas, 1870
- Allsopp, 1983
- Blackburn, 1898
- Rakovič, 1986
- Gordon & Pittino, 1992
- Thomson, 1878
- Arrow, 1908
- Branco, 1990
- Schoch, 1897
- Brenske, 1894
- Miyake & Yamaya, 1994
- Dellacasa, Dellacasa & Skelley, 2010
- Rakovič & Král, 1997
- Dellacasa, Gordon & Dellacasa, 2004
- Miyake, 1985
- Frey, 1965
- Sharp, 1873
- Erichson, 1842
- Prokofiev, 2016
- Blackburn, 1898
- Scott, 1912
- Vinson, 1946
- Martinez & Pereira, 1958
- Martinez, 1952
- Smith & Mondaca, 2015
- Brenske, 1900
- Kolbe, 1908
- Hollande & Therond, 1998
- Mulsant & Rey, 1869
- Dombrow, 2002
- Matsumoto, 2016
- Paulian, 1970
- Nimbus Mulsant & Rey, 1869
- Kraatz, 1883
- Kraatz, 1880
- Nakane, 1959
- Ochi & Kawahara, 2001
- Nomura, 1973
- Sawada, 1941
- Branco, 2010
- Britton, 1957
- Allsopp & Hutchinson, 2020
- Dellacasa & Dellacasa, 2005
- Mulsant & Rey, 1869
- Petrovitz, 1962
- Blackburn, 1898
- Heller, 1896
- Quedenfeldt, 1884
- Sharp, 1875
- (Holm & Perissinotto, 2004)
- Prokofiev, 2016
- Vaz-de-Mello, 2008
- Westwood, 1879
- Bordat, 1988
- Lacroix, 1998
- Vaz-de-mello, 2003
- Moeseneder & Hutchinson, 2012
- Brenske, 1892
- Harold, 1877
- Dellacasa, Král, Dellacasa & Bordat, 2014
- Schmidt, 1917
- Boheman, 1857
- Koshantschikov, 1916
- Fairmaire, 1897
- Gordon & Pittino, 1992
- Burmeister, 1842
- MacLeay, 1871
- Král, Rakovič & Mencl, 2015
- White, 1846
- Waterhouse, 1875
- Burmeister, 1855
- Kolbe, 1897
- Kraatz, 1894
- Schönherr, 1817
- Schein, 1958
- Burmeister, 1844
- LeConte, 1856
- Kolbe, 1891
- Dejean, 1821
- Onitis Fabricius, 1798
- Erichson, 1847
- Vaz-de-Mello, 2008
- Frolov & Vaz-de-Mello, 2015
- Erichson, 1847
- Reiche, 1860
- Onthophagus Latreille, 1802
- Arrow, 1914
- Moser, 1916
- Frey, 1972
- Boucomont, 1912
- Péringuey, 1904
- Solier, 1851
- Britton, 1987
- MacLeay, 1819
- MacLeay, 1838
- Britton, 1957
- Král, Rakovič & Mencl, 2016
- Burmeister, 1842
- Lacroix, 1998
- Kolbe, 1897
- Kolbe, 1904
- Fairmaire, 1878
- Endrödi, 1983
- Krikken, 1982
- Schmidt, 1914
- Reitter, 1900
- Mulsant & Rey, 1869
- MacLeay, 1819
- Ohaus, 1912
- Burmeister, 1847
- Carne, 1957
- Brenske, 1900
- Reitter, 1895
- Bates, 1870
- Oryctes Hellwig, 1798
- † Ratcliffe, Smith & Erwin, 2005
- Endrödi, 1978
- Boisduval, 1835
- Guérin-Méneville, 1830
- Kolbe, 1910
- Gordon & Skelley, 2007
- Branco & Baraud, 1988
- Le Peletier & Serville, 1828
- Brenske, 1896
- Mulsant, 1842
- D'Andretta & Martinez, 1957
- Scholtz & Howden, 1987
- Péringuey, 1902
- Costa, Cherman & Iannuzzi, 2020
- Frey, 1968
- Arrow, 1917
- Dellacasa & Stebnicka, 2001
- Quedenfeldt, 1888
- Solsky, 1876
- Arrow, 1908
- Dellacasa, Dellacasa & Gordon, 2016
- Dejean, 1833
- Pouillaude, 1920
- Brenske, 1899
- Oxysternon Castelnau, 1840
- Mulsant, 1842
- Skelley, 2007
- Soula, 2006
- Brenske, 1894
- Pachnoda Burmeister, 1842
- Alexis & Delpont, 2002
- Blanchard, 1851
- Brenske, 1895
- Brenske, 1903
- Le Peletier & Serville, 1828
- Erichson, 1847
- Laporte, 1832
- Reitter, 1902
- Moser, 1920
- Germar, 1848
- Griffith & Pidgeon, 1831
- Arrow, 1908
- Fairmaire, 1884
- Dejean, 1821
- Ohaus, 1912
- Brenske, 1897
- MacLeay, 1821
- Blanchard, 1851
- De Palma & Malec, 2020
- Hope, 1841
- Curtis, 1845
- Lacroix, 1999
- Kolbe, 1910
- Reiche, 1871
- Thomson, 1880
- † Poinar, 2011
- Stebnicka, 1985
- Lewis, 1895
- Kraatz, 1880
- Reitter, 1902
- Burmeister, 1847
- Burmeister, 1842
- Papuana Arrow, 1911
- Hollande & Thérond, 1998
- Ohaus, 1915
- Péringuey, 1904
- Balthasar, 1938
- Dombrow, 2002
- Burmeister, 1842
- Soula, 2002
- Harold, 1873
- Bates, 1888
- Waterhouse, 1875
- Balthasar, 1963
- Wasmann, 1918
- Ohaus, 1915
- Balthasar, 1932
- Howden & Cook, 2002
- Howden, 1971
- Landin, 1959
- Balthasar, 1937
- Bordat, 2022
- Machatschke, 1957
- Soula, 1998
- Vulcano, Pereira & Martinez, 1966
- Becker, 1910
- Shipp, 1897
- Moser, 1924
- Morón, 1983
- Lacroix & Coache, 2017
- Roggero, Barbero & Palestrini, 2015
- Antoine, 1989
- Ocampo & Colby, 2009
- Ohaus, 1915
- Nomura, 1974
- Lacroix, 1998
- Seidlitz, 1888
- Lacroix, 1997
- Antoine, 1988
- Schmidt, 1914
- Cockerell, 1905
- Casey, 1915
- Martinez, 1961
- Balthasar, 1969
- Endrödi, 1978
- Harold, 1877
- Arrow, 1910
- Mikšić, 1985
- Chalumeau, 1992
- Hauser, 1904
- Moser, 1907
- Verdu, Stebnicka & Galante, 2006
- Soula, 2002
- Prell, 1934
- Kraatz, 1899
- Blackburn, 1911
- Paulian, 1991
- Balthasar, 1955
- Paulian, 1932
- Ohaus, 1904
- Britton, 1990
- Reitter, 1896
- Westwood, 1842
- Antoine, 2005
- Pacheco, Monné, Vaz-de-Mello & Ahrens, 2022
- Balthasar, 1961
- Ohaus, 1915
- Shipp, 1895
- Ohaus, 1915
- Bordat, 2013
- Janson, 1881
- Baraud, 1962
- Lacroix, Coache & Filippi, 2022
- Holm, 1988
- Gordon & Skelley, 2007
- Marais & Holm, 1989
- Kraatz, 1893
- Moser, 1918
- Prell, 1934
- Kraatz, 1880
- Ohaus, 1915
- Ohaus, 1915
- Bates, 1888
- Schaufuss, 1890
- Balthasar, 1963
- Paulian, 1984
- Britton, 1990
- Fairmaire, 1903
- Dechambre, 1994
- Brenske, 1899
- Deschodt & Scholtz, 2008
- Péringuey, 1904
- Balthasar, 1945
- Soula, 2008
- Balthasar, 1938
- Martinez & D'andretta, 1956
- Scholtz & Howden, 1987
- Ballion, 1871
- Arrow, 1913
- Laporte, 1832
- Harold, 1868
- Kraatz, 1880
- Matsumoto, 2016
- Erichson, 1847
- Pelidnota MacLeay, 1819
- † Cockerell, 1920
- Burmeister, 1847
- Paulian, 1985
- Moser, 1926
- Ohaus, 1919
- De Wailly, 1950
- Brenske, 1896
- Brenske, 1891
- Kraatz, 1892
- Endrödi, 1968
- Hope, 1837
- Péringuey, 1901
- Krikken, 1983
- Westwood, 1847
- Vaz-de-Mello, 2008
- Péringuey, 1904
- Brenske, 1896
- Burmeister, 1847
- Kraatz, 1880
- Lacroix, 2006
- Kolbe, 1910
- Brenske, 1898
- Waterhouse, 1875
- Burmeister, 1844
- Lacroix, 1997
- Mondaca & Valencia, 2016
- Blackburn, 1898
- Mikšić, 1965
- Rakovič, 1979
- Reitter, 1903
- Reitter, 1892
- Waterhouse, 1841
- Paulian, 1985
- Hope, 1841
- Hope, 1833
- Motschulsky, 1859
- Burmeister, 1844
- Reitter, 1889
- Quedenfeldt, 1884
- Erichson, 1848
- MacLeay, 1819
- Kraatz, 1894
- Blanchard, 1851
- Reitter, 1892
- Brenske, 1897
- Heller, 1900
- Dechambre, 2008
- Latreille, 1807
- Zídek, 2021
- MacLeay, 1838
- Dejean, 1833
- Brèthes, 1919
- Brenske, 1897
- LeConte, 1856
- Bordat, 2009
- Kolbe, 1894
- Moser, 1919
- Kraatz, 1880
- Britton, 1987
- Brenske, 1897
- Schaum, 1844
- Schoch, 1898
- Broun, 1886
- Blackburn, 1890
- Moser, 1918
- Blanchard, 1846
- Waterhouse, 1876
- Eschscholtz, 1830
- Stephens, 1830
- Phyllophaga Harris, 1826
- Brenske, 1899
- Blackburn, 1898
- Fischer Von Waldheim, 1823
- Brenske, 1900
- Blanchard, 1851
- Soula, 2002
- Bourgoin, 1921
- Kraatz, 1888
- Burmeister, 1842
- Moser, 1924
- Erichson, 1842
- Branco, 1991
- Harold, 1875
- Ratcliffe & Giraldo, 2014
- Rakovič & Král, 1997
- Péringuey, 1900
- Brenske, 1899
- Mulsant, 1842
- Dellacasa & Dellacasa, 2005
- Dellacasa & Dellacasa, 2005
- Mulsant & Rey, 1869
- Chapin, 1932
- Burmeister, 1844
- Dejean, 1833
- Schmidt, 1917
- MacLeay, 1819
- Krikken, 1978
- Westwood, 1854
- Janssens, 1942
- Ohaus, 1910
- Bates, 1888
- Westwood, 1874
- Mulsant, 1842
- Le Peletier & Serville, 1828
- Horn, 1867
- Wallace, 1867
- Péringuey, 1904
- Erichson, 1847
- Lacroix & Montreuil, 2019
- Lacroix, 2006
- Jameson, 1997
- Morón, 1983
- Schmidt, 1914
- Lansberge, 1875
- Kraatz, 1892
- Mulsant, 1842
- Fairmaire, 1896
- Ruter, 1978
- Brenske, 1899
- Brenske, 1899
- Burmeister, 1847
- Burmeister, 1847
- Harold, 1869
- Moser, 1915
- Moser, 1917
- Howden, 1997
- Paulian, 1942
- Schmidt, 1914
- Arrow, 1910
- Poecilopharis Kraatz, 1880
- Kraatz, 1880
- Arrow, 1910
- Burmeister, 1842
- Waterhouse, 1875
- De Wailly, 1950
- Polyphylla Harris, 1841
- Blanchard, 1851
- Janson, 1888
- Kraatz, 1882
- Popillia Dejean, 1821
- Burmeister, 1842
- Arrow, 1911
- Krikken, 1984
- † Skelley, 2021
- Jákl, 2018
- Harold, 1869
- Lansberge, 1883
- Reitter, 1903
- Blanchard, 1851
- Deyrolle, 1864
- Britton, 1988
- Erichson, 1847
- Harold, 1861
- López-García & Deloya, 2022
- Broun, 1904
- Brenske, 1893
- Sigwalt, 1987
- † Bai, Ren & Yang, 2011
- Newman, 1836
- Kolbe, 1905
- Protaetia Burmeister, 1842
- Mikšić, 1968
- Britton, 1987
- Brenske, 1896
- Krikken, 1976
- Krikken, 1978
- Decelle, 1979
- † Nikolajev, 2007
- † Nikolajev, 2000
- † Nikolajev, Wang & Zhang, 2014
- Endrödi, 1976
- Fallen, 1807
- Thomson, 1859
- Stebnicka, 1994
- Brenske, 1899
- Péringuey, 1904
- Reitter, 1892
- Semenov, 1890
- Heller, 1923
- Schmidt, 1914
- Antoine, 1989
- Bourgoin, 1921
- Brown, 1927
- Fairmaire, 1901
- Lacroix, 1991
- Kraatz, 1880
- Hollande & Thérond, 1998
- Orbigny, 1896
- Schein, 1959
- Blackburn, 1911
- Paulian, 1984
- Kraatz, 1880
- Frey, 1971
- Bordat, 1994
- Dellacasa & Dellacasa, 1997
- Soula, 1998
- Bordat, 2016
- Keith, 2000
- Bates, 1887
- Blackburn, 1890
- Ritsema, 1882
- Mikšić, 1985
- Ferreira, 1977
- Ohaus, 1905
- Kraatz, 1882
- Neervoort Van De Poll, 1886
- Dellacasa, Dellacasa & Gordon, 2013
- Moser, 1924
- Ferreira, 1960
- Hardy, 1971
- Frolov & Montreuil, 2019
- Mikšić, 1974
- Gutiérrez, 1950
- Nonfried, 1894
- Soula, 1998
- (Bordat, 1993)
- Ohaus, 1910
- Dellacasa, Gordon & Dellacasa, 2007
- Dellacasa & Dellacasa, 2000
- Blackburn, 1892
- Arrow, 1908
- Soula, 1998
- Moser, 1919
- Ohaus, 1903
- Nomura, 1974
- Ohaus, 1912
- Lacroix, 1998
- Pittino, 1984
- Minkina, 2023
- Paulian, 1984
- Baraud, 1985
- Balthasar, 1930
- Petrovitz, 1969
- Lacroix, 1997
- Katovich, 2011
- Felsche, 1904
- Keith & Reichenbach, 2018
- Bordat, 1990
- Duvivier, 1892
- Moser, 1918
- Dellacasa, 1990
- Soula, 2002
- Benderitter, 1913
- Sharp, 1873
- Balthasar, 1941
- Krikken, 1976
- Ohaus, 1904
- Keith, 2006
- Guérin-Méneville, 1830
- Kobayashi, 1980
- Ricchiardi, 2015
- Bordat & Howden, 1995
- Dalle Torre, 1912
- Péringuey, 1901
- Kraatz, 1882
- Dellacasa, Dellacasa & Gordon, 2015
- Dellacasa, 1986
- Soula, 2002
- Keith, 2008
- Mikšić, 1967
- Jacquelin Du Val, 1859
- Keith, 2007
- Quedenfeldt, 1884
- Schein, 1959
- Petrovitz, 1962
- Péringuey, 1904
- Burmeister, 1842
- Broun, 1895
- Burmeister, 1855
- Brenske, 1892
- Bates, 1888
- Murayama, 2010
- Kraatz, 1883
- Redtenbacher, 1868
- Mulsant & Rey, 1869
- Ohaus, 1910
- Ocampo & Smith, 2006
- Muramoto, 2006
- Smith, 2008
- Arrow, 1931
- Thomson, 1859
- Burmeister, 1842
- Kolbe, 1884
- Boisduval, 1835
- Pyrrhopoda Kraatz, 1880
- Král, 1997
- Péringuey, 1902
- Thomson, 1877
- Pittino, 2006
- Kolbe, 1894
- Lacroix, 1991
- Saylor, 1947
- Schein, 1958
- Fairmaire, 1860
- Reineria Mikšić, 1968
- Lacroix, 1993
- Balthasar, 1951
- Frolov & Montreuil, 2009
- MacLeay, 1819
- Burmeister, 1855
- Burmeister, 1842
- Krikken, 1981
- Lacroix, 1997
- Arrow, 1930
- Kraatz, 1900
- Burmeister, 1842
- Ohaus, 1912
- Klug, 1855
- Ruter, 1965
- Perty, 1833
- Maruyama, 2009
- Burmeister, 1842
- Burmeister, 1844
- Ohaus, 1909
- Péringuey, 1901
- Westwood, 1841
- Kolbe, 1894
- Rhizotrogus Latreille, 1825
- Adam, 1994
- Hope, 1837
- Erichson, 1847
- Ohaus, 1913
- Lacroix, 2008
- Fairmaire, 1883
- Burgeon, 1942
- Agassiz, 1846
- Bedel, 1911
- Reitter, 1892
- Clouët, 1900
- Mulsant, 1842
- Moser, 1921
- Moser, 1924
- Burmeister, 1842
- Hutchinson & Moeseneder, 2022
- Lacroix, 1997
- Lacroix, 1996
- De Wailly, 1950
- Matsumoto, 2016
- Gordon & Skelley, 2007
- Machatschke, 1957
- Latreille, 1802
- Waterhouse, 1874
- Bates, 1888
- Arrow, 1908
- Krikken, 1980
- Montreuil & Lacroix, 2020
- Lacroix, 1998
- Morón & Grossi, 2015
- Dellacasa & Dellacasa, 2000
- Brookes, 1944
- Sharp, 1873
- Stebnicka & Galante, 2007
- Balthasar, 1967
- Redtenbacher, 1857
- Stebnicka, 1993
- Blackburn, 1904
- Rakovič & Mencl, 2011
- Erichson, 1847
- Blackburn, 1907
- Waterhouse, 1878
- Paulian, 1934
- Frey, 1975
- Gordon & Skelley, 2007
- Burmeister, 1847
- Evans, 1987
- Quedenfeldt, 1884
- Ohaus, 1912
- Schaum, 1841
- Péringuey, 1901
- Balthasar, 1965
- Scarabaeus Linnaeus, 1758
- Erichson, 1847
- Erichson, 1835
- Génier & Kohlmann, 2003
- Westwood, 1837
- Dombrow, 2001
- Burmeister, 1844
- Gordon & Skelley, 2007
- Ruter, 1957
- Gautier Des Cottes, 1872
- Arrow, 1917
- Blanchard, 1850
- Fischer Von Waldheim, 1823
- Dejean, 1833
- Kirby, 1825
- Berg, 1898
- Erichson, 1842
- Britton, 1990
- Blackburn, 1892
- Martinez, 1948
- Martinez, 1953
- Broun, 1886
- Kraatz, 1880
- Péringuey, 1904
- Castelnau, 1840
- Brenske, 1897
- Fairmaire, 1891
- Stebnicka, 1999
- Hope, 1847
- De Wailly, 1950
- Serica MacLeay, 1819
- Motschulsky, 1860
- Boisduval, 1835
- Guérin-Méneville, 1839
- Sharp, 1882
- Kabakov, 1996
- Miyake, 2002
- Gordon & Skelley, 2007
- Stebnicka, 1994
- Masumoto, 1991
- Keith, 2005
- Reitter, 1896
- Mulsant & Rey, 1869
- Landin, 1949
- Roets & Oberlander, 2010
- Vaz-de-Mello, 2008
- Harold, 1871
- Kraatz, 1886
- Červenka, 1994
- Boucomont, 1928
- Blanchard, 1851
- Dellacasa, 1986
- Simonis, 1985
- Dechambre, 1998
- Sisyphus Latreille, 1807
- Fairmaire, 1894
- Dellacasa, Dellacasa, Gordon & Stebnicka, 2011
- Westwood, 1847
- Král, Sehnal & Bezděk, 2012
- Lansberge, 1882
- Brenske, 1899
- Fairmaire, 1887
- Soula, 2006
- Kolbe, 1894
- Castelnau, 1840
- Arrow, 1902
- Lansberge, 1886
- Kakizoe, Jiang & Wang, 2021
- Jameson, 1997
- Burmeister, 1855
- Brenske, 1900
- Brenske, 1899
- Kolbe, 1904
- Kraatz, 1890
- Sharp, 1877
- Westwood, 1848
- Kraatz, 1892
- Ohaus, 1910
- Machatschke, 1970
- Burmeister, 1847
- Burmeister, 1855
- Burmeister, 1842
- Skelley, 2007
- Burmeister & Schaum, 1840
- Britton, 1990
- Burmeister, 1847
- Fairmaire, 1889
- Arrow, 1943
- Moser, 1913
- Brenske, 1900
- Nomura, 1977
- Karsch, 1881
- Burmeister, 1842
- Schmidt, 1914
- Brenske, 1896
- Stephanorrhina Burmeister, 1842
- Arrow, 1905
- Hope, 1837
- Bainbridge, 1840
- Valck Lucassen, 1933
- Dombrow, 2001
- Kolbe, 1914
- Branco, 1989
- Harold, 1871
- Branco, 1989
- Kolbe, 1897
- Hasenpusch & Moeseneder, 2010
- Fairmaire, 1901
- Strategus Hope, 1837
- Lansberge, 1874
- Burmeister, 1844
- Gordon & Skelley, 2007
- Gory & Percheron, 1833
- Ohaus, 1912
- Mulsant & Rey, 1869
- Reitter, 1894
- Branco, 1992
- Olsoufieff, 1924
- Péringuey, 1904
- Mencl, 2011
- Martinez, 1955
- Paulian, 1954
- Fairmaire, 1896
- Boheman, 1857
- Halffter & Martinez, 1977
- Erichson, 1835
- Schmidt, 1914
- Fairmaire, 1899
- Bates, 1868
- Britton, 1956
- Kolbe, 1897
- Lacroix, 1993
- Brenske, 1898
- Kolbe, 1897
- Péringuey, 1901
- Péringuey, 1901
- Janssens, 1942
- Sharp, 1877
- Kraatz, 1895
- Burmeister, 1842
- Valck Lucassens, 1939
- Kobayashi, 1990
- Yamaya, 2001
- Brenske, 1899
- Stebnicka, 2006
- Petrovitz, 1971
- Ménétriés, 1832
- Lacroix, 2008
- Lacroix, 2002
- Scholtz & Howden, 1987
- Quedenfeldt, 1888
- Thomson, 1880
- Moser, 1924
- Miyake, 1993
- Burmeister, 1842
- Endrödi, 1975
- Sharp, 1873
- Burmeister, 1844
- Krikken, 1975
- Erichson, 1842
- Britton, 1990
- Westwood, 1841
- Hope, 1837
- Burmeister, 1842
- Brenske, 1900
- Brenske, 1897
- Lacroix, 1989
- Krikken, 1970
- Mateu, 1966
- Nakane, 1961
- Wasmann, 1894
- Britton, 1957
- Wasmann, 1918
- Reichensperger, 1915
- Krikken, 2018
- Rakovič, 1981
- Hope, 1837
- Vaz-de-Mello & Halffter, 2006
- Matsumoto, 2021
- Schmidt, 1914
- Blanchard, 1845
- Kraatz, 1896
- Duarte & Grossi, 2022
- Blanchard, 1842
- Ahrens, 2004
- Mulsant, 1842
- Péringuey, 1902
- Thomson, 1880
- Huchet, 2000
- Soula, 2006
- Semenov & Reichardt, 1925
- Prokofiev, 2015
- Decelle, 1968
- Burmeister, 1847
- Brenske, 1897
- LeConte, 1856
- Blackburn, 1904
- Reitter, 1898
- Burmeister, 1844
- Ohaus, 1915
- Prokofiev, 2014
- Burmeister, 1842
- Roggero, Moretto, Barbero & Palestrini, 2019
- Krikken, 2009
- Miyake, 1994
- Krikken, 1978
- Péringuey, 1901
- Soula, 2002
- Péringuey, 1904
- Westwood, 1841
- Reitter, 1902
- Erichson, 1847
- Orbigny, 1904
- Prokofiev, 2021
- Arrow, 1907
- Pavicevic, 1985
- Dombrow, 2002
- Fairmaire, 1878
- Brenske, 1899
- Tragiscus Klug, 1855
- Faldermann, 1835
- Kolbe, 1914
- Dupuis & Perrin, 2022
- Reitter, 1903
- Arrow, 1917
- Schmidt, 1914
- Paulian, 1942
- Schmidt, 1914
- Kraatz, 1880
- Erichson, 1847
- Vaz-de-Mello, 2008
- Harold, 1868
- Waterhouse, 1875
- Minkina, 2018
- Burmeister, 1855
- Petrovitz, 1963
- Clouët, 1901
- Trichiotinus Casey, 1915
- Trichius Fabricius, 1775
- Moser, 1916
- Burmeister, 1847
- Blanchard, 1851
- Nomura, 1952
- Nomura, 1974
- Legrand, 2001
- Balthasar, 1945
- Bedel, 1911
- Bordat, 2022
- Burmeister, 1842
- Ohaus, 1897
- Moser, 1917
- Burmeister, 1842
- Brenske, 1896
- Kraatz, 1894
- Burmeister & Schaum, 1840
- Pouillaude, 1913
- Hope, 1831
- (Petrovitz, 1963)
- Brenske, 1893
- Brenske, 1894
- Reitter, 1919
- Moser, 1921
- Westwood, 1845
- Saylor, 1942
- Ohaus, 1904
- Burmeister, 1847
- Moser, 1922
- Krikken, 1987
- Burmeister, 1847
- Balthasar, 1932
- Ohaus, 1917
- Reitter, 1896
- Brenske, 1900
- Laporte, 1832
- Boheman, 1857
- Janssens, 1937
- Mulsant, 1842
- Blanchard, 1851
- Gerstaecker, 1867
- Minck, 1920
- Lacroix, 1997
- Péringuey, 1904
- Semenov, 1905
- Ochi & Kon, 2017
- Saylor, 1945
- Burmeister, 1842
- Josso & Prévost, 2008
- Deschodt, Sole & Scholtz, 2020
- Westwood, 1842
- De Wailly, 1950
- Ruter, 1973
- Fairmaire, 1899
- Valgus Scriba, 1790
- Schein, 1958
- Lacroix, 1993
- Ohaus, 1915
- Deschodt, Davis & Scholtz, 2011
- Lacroix, 1993
- Ruter, 1964
- Jameson, 1990
- Dellacasa, Dellacasa & Bordat, 2002
- Volinus Mulsant & Rey, 1869
- Itoh, 1993
- Allsopp, Jákl & Rey, 2023
- Cambefort, 1977
- Allsopp, 1988
- Smith & Evans, 2005
- Britton, 1995
- Britton, 1988
- Dechambre, 1999
- Reitter, 1902
- Kraatz, 1881
- Arrow, 1915
- Martinez, 1952
- Arrow, 1920
- Soula, 2002
- Brême, 1844
- Villatoro & Jameson, 2001
- Hinton, 1934
- Bates, 1889
- Bates, 1904
- Kolbe, 1896
- Arrow, 1910
- Ahrens, 2005
- Moser, 1917
- Britton, 1978
- Saylor, 1937
- Harold, 1869
- Burmeister, 1842
- Holm, 1992
- Boisduval, 1835
- Hope, 1837
- Prell, 1934
- Xylotrupes Hope, 1837
- Britton, 1990
- Britton, 1987
- Britton, 1987
- Britton, 1987
- Morón & Nogueira, 1998
- Nomura, 1952
- † Nikolajev, 2015
- Bordat, 1988
- Ochi & Kon, 1996
- Saylor, 1945
- Fuhrmann, 2021
- Fairmaire, 1894
- Blackburn, 1894
- Lacroix, 1997
- Lacroix, 2006
- Arrow, 1932
Les scarabées dans la culture
Le Scarabée sacré (Scarabaeus sacer) est un animal vénéré dans l'Égypte antique. Symbole du soleil et de Rê, il était l'un des douze animaux sacrés associé aux douze heures du jour et de la nuit[réf. nécessaire].
Le Scarabée rhinocéros siamois ou scarabée de combat (Xylotrupes gideon) est capturé dans la nature puis dressé pour des duels qui font l'objet de paris dans le nord de la Thaïlande. Ces duels de scarabées sont couramment organisés. Ces combats d’insectes du type xylotrupes appelés Kwang localement sont à bien des égards surprenants, notamment du fait qu’il est sans doute plus difficile d’attribuer à un scarabée qu’à un autre animal des caractéristiques anthropomorphiques et qu'il est plus compliqué encore de deviner ce qu’il perçoit. Le duel de scarabées met à l’épreuve une forme de zoomanité trouble fondée sur une forme d’attachement inédite entre l’homme et l’animal,.
Notes et références
- Museum de l'université du Nebraska
- Michael Chinery, Insectes de France et d'Europe occidentale, Flammarion, 2012 (1re éd. 1986), p. 264
- Anatole Bailly ; 2020 : Hugo Chávez, Gérard Gréco, André Charbonnet, Mark De Wilde, Bernard Maréchal & contributeurs, « Le Bailly », (consulté le ).
- Informations lexicographiques et étymologiques de « Scarabée » dans le Trésor de la langue française informatisé, sur le site du Centre national de ressources textuelles et lexicales
- Latreille, 1802: Histoire naturelle, générale et particulière des Crustacés et des Insectes. Tome 3. F. Dufart, Paris. Internet Archive BHL
- Catalogue of Life Checklist, consulté le 23 juin 2023.
- Stéphane Rennesson, Emmanuel Grimaud, Nicolas Cesard, Duel en miniature. La délicate mise en scène des combats de scarabées dans le nord de la Thaïlande , Insectes, 151 (4), 3-6,2008
- Thiti Wannamontha, « Jeux. Le duel de scarabées, une passion thaïlandaise », sur courrierinternational.com, Courrier international,
Voir aussi
Articles connexes
- Scarabée du taro
Liens externes
- (en) Animal Diversity Web : Scarabaeidae (consulté le )
- (en) Catalogue of Life : Scarabaeidae Latreille, 1802 (consulté le )
- (fr + en) EOL : Scarabaeidae (consulté le )
- (en) Fauna Europaea : Scarabaeidae (consulté le )
- (fr + en) GBIF : Scarabaeidae (consulté le )
- (fr) INPN : Scarabaeidae Latreille, 1802 (TAXREF) (consulté le )
- (en) IRMNG : Scarabaeidae Latreille, 1802 (consulté le )
- (fr + en) ITIS : Scarabaeidae Latreille, 1802 (consulté le )
- (en) NCBI : Scarabaeidae (taxons inclus) (consulté le )
- (en) OEPP : Scarabaeidae (consulté le )
- (en) Paleobiology Database : Scarabaeidae (consulté le )
- (en) Taxonomicon : Scarabaeidae Latreille, 1802 (consulté le )
- (en) Tree of Life Web Project : Scarabaeidae (consulté le )
- (en) UICN : taxon Scarabaeidae (consulté le )
- (en) WoRMS : Scarabaeidae Latreille, 1802 (+ liste espèces) (consulté le )
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Portail de l’entomologie
Auteur: www.NiNa.Az
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Scarabee redirige ici Pour les autres significations voir Scarabee homonymie Ne doit pas etre confondu avec Scarabaeoidea Les Scarabaeidae en francais Scarabeides sont une famille d insectes coleopteres de la super famille des Scarabaeoidea Ils sont connus sous le nom vernaculaire de Scarabee C est notamment la famille des dynastes du scarabee rhinoceros europeen des bousiers et des hannetons Scarabaeidae Accouplement de Classification ITISRegne AnimaliaSous regne BilateriaInfra regne ProtostomiaSuper division EcdysozoaEmbranchement ArthropodaSous embr HexapodaClasse InsectaSous classe PterygotaInfra classe NeopteraSuper ordre HolometabolaOrdre ColeopteraSous ordre PolyphagaInfra ordre ScarabeiformiaSuper famille Scarabaeoidea FamilleScarabaeidae Latreille 1802 Les limites de cette famille sont encore debattues et controversees Les chercheurs europeens en ont une definition plutot restrictive alors qu un certain nombre de chercheurs americains y incorporent bien plus de taxons Les antennes des insectes de cette famille sont en massue dont l extremite peut s ouvrir en un eventail de feuillets L extremite de l abdomen est decouverte en raison d elytres normalement tronques La plupart des especes volent correctement Plusieurs de ces especes stridulent en frottant l extremite de leurs elytres contre la face dorsale de leur abdomen Cette vaste famille comprend des coprophages et des saproxylophages nuisibles aux vegetaux Noms vernaculairesLe terme de Scarabee provient via le vieux francais escarbot du latin scarabaeus par alteration du grec ancien karabos karabos escarbot scarabee insecte egalement a l origine du terme carabe Le terme scarabee designe de facon generale en zoologie l ensemble de la super famille des Scarabeoides mais ce terme peut designer chez certains auteurs d une facon plus restrictive la famille des Scarabaeidae et meme pour certains uniquement les coprophages de cette famille ref necessaire Plusieurs groupes qui peuvent etre paraphyletiques de ces insectes ont des noms vernaculaires tels que les Dynastes ou scarabees rhinoceros les bousiers les cetoines les hannetons Certaines especes ont un nom vernaculaire particulier comme le Pique prune Osmoderma eremita SystematiqueLa famille a ete decrite par l entomologiste francais Pierre Andre Latreille en 1802 Liste des sous familles Selon le Catalogue of Life 23 juin 2023 cette famille se divise en 28 sous familles Aclopinae Blanchard 1850 Aegialiinae Castelnau 1840 Arrow 1940 Aphodiinae Leach 1815 Janssens 1946 Cetoniinae Leach 1815 Blanchard 1845 Nikolajev 1995 Arrow 1911 Dynastinae MacLeay 1819 Stebnicka 1977 Latreille 1829 Burmeister 1844 Nikolajev 1992 Melolonthinae MacLeay 1819 LeConte 1861 Orphninae Erichson 1847 Reitter 1902 Erichson 1847 Howden 1997 Nikolajev 2000 Rutelinae MacLeay 1819 Scarabaeinae Latreille 1802 Sericinae Kirby 1837 Erichson 1847 Wasmann 1918 Trichiinae Kolbe 1897 Valginae Schenkling 1922 Liste des genres Selon le Catalogue of Life 23 juin 2023 cette famille regroupe 2 590 genres Dejean 1833 Blanchard 1850 Schmidt 1909 Acanthobodilus Dellacasa 1983 Janssens 1937 Kraatz 1895 Prell 1936 Britton 1988 Erichson 1848 Erichson 1840 Kraatz 1888 Erichson 1835 Casey 1889 Arrow 1899 Ohaus 1912 Schmidt 1914 Schmidt 1914 Acrossus Mulsant 1842 Kraatz 1882 Westwood 1841 Ohaus 1909 Bordat amp Howden 1995 Ratcliffe 2014 Schmidt 1914 Kraatz 1883 Blanchard 1851 Adoretus Dejean 1833 Brenske 1893 Brenske 1893 Arrow 1917 Blackburn 1889 Britton 1995 Villiers 1956 Latreille 1807 Paulian 1984 Arrow 1904 Westwood 1845 Burmeister 1847 Dejean 1833 Arrow 1899 Decelle 1982 Burmeister 1842 Petrovitz 1969 Dellacasa 1984 Pittino amp Mariani 1986 Krikken 2009 Bordat 2019 Decelle 1968 Bordat 2016 Bates 1887 Reitter 1895 Le Peletier amp Serville 1828 Moser 1918 Eschscholtz 1829 Brenske 1896 Miyake amp Iwase 1991 Schmidt 1914 Mulsant amp Rey 1869 Dellacasa Dellacasa amp Gordon 2008 Agrilinus Mulsant amp Rey 1869 Baguena 1930 Balthasar 1963 Stebnicka amp Howden 1996 Kral Mencl amp Rakovic 2021 Petrovitz 1980 Allsopp 2018 Lacroix 2003 Arrow 1908 Deschodt amp Scholtz 2008 Britton 1955 Schenkling 1921 Arrow 1940 Bordat 2009 Petrovitz 1963 Janssens 1940 Peringuey 1904 Allomyrina Arrow 1911 Janssens 1936 Dupuis amp Dechambre 2001 Fauvel 1903 Burmeister 1842 Boucomont 1923 Britton 1978 Schmidt 1914 Peringuey 1904 Frey 1975 Blanchard 1850 Reitter 1902 Kolbe 1895 Kraatz 1882 Blanchard 1851 Reitter 1902 Dombrow 2002 Westwood 1878 Kolbe 1910 MacLeay 1819 Skelley 2007 Stebnicka amp Skelley 2003 Amidorus Mulsant 1870 Cambefort 1981 Nomura 1974 Thomson 1878 Bordat 1999 Mulsant 1842 Fairmaire 1898 Huchet 2000 Amphimallon Latreille 1825 Lansberge 1874 Brenske 1894 Smith 2008 Dechambre amp Madge 1979 Blanchard 1842 Britton 1990 Petrovitz 1969 Arrow 1922 Martinez 1958 Antoine 1989 Lacroix 1995 Kolbe 1894 Breme 1844 Burmeister 1842 Peringuey 1907 Curtis 1835 Erichson 1847 Britton 1957 Kolbe 1892 Lacroix 1993 Blackburn 1898 Martinez amp Pereira 1956 Burmeister 1844 Latreille 1807 Moser 1913 Fischer Von Waldheim 1824 Westwood 1842 Paulian 1954 Howden 1971 Burmeister 1842 Moser 1914 Sharp 1875 Ohaus 1901 Fairmaire 1878 Anomala Samouelle 1819 Fric 1885 Moser 1913 Arrow 1908 Reitter 1887 Jameson Paucar Cabrera amp Solis 2003 Paulian 1952 Blackwelder 1944 Westwood 1842 Arrow 1946 Mulsant amp Rey 1869 Fairmaire 1897 Saylor 1940 Janssens 1950 Arrow 1917 MacLeay 1838 Simonis 1981 Rivers 1889 Anoplognathus Leach 1815 Brulle 1837 Arrow 1939 Laporte 1832 Petrovitz 1971 Lacroix 1993 Minkina 2020 Gomez Pallerola 1979 Medvedev 1949 Britton 1957 Burmeister 1842 Moser 1918 Lacroix 1993 Eschscholtz 1818 Soula 1998 Lu Ahrens Bai Shih amp Ren 2023 Brenske 1900 Burmeister 1855 Breme 1844 Petrovitz 1958 Howden 1968 Brenske 1900 Britton 1990 Kraatz 1880 Antoine 1987 Harold 1868 Peringuey 1901 Koshantschikov 1911 Schmidt 1914 Paulian 1942 Aphodius Hellwig 1798 Peringuey 1901 Endrodi 1983 Endrodi 1964 Rivers 1889 Kolbe 1905 LeConte 1856 Cartwright 1952 Lacroix 1993 Blanchard 1850 Antoine 2001 Kirby 1818 Lansberge 1886 Brenske 1903 Olsoufieff 1947 Baraud 1962 Schmidt 1917 Matthews 1974 Montreuil 2011 Smith amp Evans 2018 Gutierrez 1952 Janssens 1937 Uliana amp Sabatinelli 2017 Westwood 1842 Moron amp Krikken 1990 Nikolajev 1982 Minkina 2020 Brenske 1895 Kolbe 1905 Kolbe 1910 Brenske 1897 Burmeister 1844 Paulsen 2021 MacLeay 1819 Penco amp Zubaran 2013 Stebnicka amp Dellacasa 2003 Dombrow 2003 Thomson 1878 Rossi 1958 Peringuey 1904 Martinez 1960 Stebnicka 1999 Brenske 1894 Brancsik 1893 Makhan 2006 Nikolajev 2000 Medvedev 1949 Petrovitz 1961 Bates 1888 Rojkoff 2013 Aspilus Westwood 1874 Erichson 1847 Martinez 1957 Brenske 1898 Westwood 1874 Westwood 1843 Britton 1990 Peringuey 1907 Schurhoff 1937 Petrovitz 1973 Ataenius Harold 1867 Petrovitz 1954 Bedel 1892 Meunier 1898 Ateuchus Weber 1801 Lacroix 1998 Erichson 1835 Gonzalez Alvarado Molano Rendon amp Vaz de Mello 2019 Kraatz 1898 Distant 1911 Philips amp Bell 2008 Strand 1942 Burmeister 1847 Burmeister 1841 Guerin Meneville 1838 White 1859 Decelle 1973 Brenske 1900 Klug 1838 Hutchinson amp Moeseneder 2019 Deschodt amp Davis 2018 Mencl 2008 Petrovitz 1958 Balthasar 1942 Howden amp Storey 1992 Britton 1978 Brenske 1897 Krikken 2018 Miksic 1976 Thomson 1880 Machatschke 1970 Thomson 1877 Nikolajev 2005 Paucar cabrera 2003 Fairmaire 1898 Gordon amp Skelley 2007 Paulian 1986 Thomson 1880 Allsopp 2022 Ratcliffe 1981 Blanchard 1850 Endrodi 1956 Schein 1957 Chalumeau 1983 Pereira amp Vulcano amp Martinez 1960 Harold 1869 Dombrow 2005 Peringuey 1904 Vaz de Mello 2008 Fairmaire 1898 Fairmaire 1893 Machatschke 1970 White 1841 Mulsant amp Rey 1869 Antoine 2003 Lacroix 1993 Paulian 1961 Peringuey 1902 Fairmaire 1898 Schmidt 1914 Burmeister 1842 Lacroix 1998 Lacroix 1994 Blanchard 1850 Peringuey 1902 Reitter 1903 Endrodi 1973 Dellacasa amp Dellacasa 2005 Adam 1994 Mulsant amp Rey 1869 Paulian 1975 Fisher Von Waldheim 1829 Harold 1868 Matthews 1974 Westwood 1848 Ruter 1978 Pittino amp Mariani 1986 Halffter 1958 Senyuz 2017 Burmeister 1842 Hope 1837 Balthasar 1932 Kraatz 1890 Koshantschikov 1913 Kolbe 1894 Brenske 1896 Kraatz 1896 Thomson 1858 Brenske 1898 Waterhouse 1881 Guerin Meneville 1831 Vaz de Mello 2008 Blanchard 1851 Baraud 1986 Berg 1898 Ricchiardi 2017 Burmeister 1842 Landin 1960 Ozdikmen amp Demir 2008 Martinez 1952 Howden amp Storey 1992 Dejan 1833 Sabatinelli amp Pontuale 1998 Nomura 1970 Berg 1898 Dombrow 2003 Miksic 1955 Harold 1869 Harold 1869 Blackburn 1892 Brenske 1898 Burmeister 1844 Balthasar 1964 Caccobius Thomson 1859 Paulian 1984 Lewis 1895 Hope 1841 Lacroix 1998 Motschulsky 1859 Krell 2011 Calicnemis Laporte 1832 Gordon amp Skelley 2007 Blackburn 1895 Lacroix 1998 Burmeister 1842 Blanchard 1851 Peringuey 1901 Blanchard 1851 Ohaus 1903 White 1845 Moser 1916 Brenske 1894 Mohnike 1873 Dellacasa 1984 Blanchard 1850 Bates 1888 Bourgoin 1917 Blanchard 1850 Howden 1970 Ricchiardi 2000 Paulian 1986 Branco 1989 Erichson 1847 Brenske 1900 Lacroix 2005 Lacroix 2002 Hope 1831 Kirby 1827 Decelle 1979 Brenske 1900 Stebnicka amp Howden 1995 Saylor 1938 Erichson 1847 Chapin 1934 Davis Scholtz amp Harrison 1999 Canthon Hoffmannsegg 1817 Chapin 1930 Paulian 1939 MacLeay 1871 Paulian 1939 Bordat 2022 Lacroix 1994 Lacroix 1999 Tesar 1945 Bezdek 2016 Endrodi 1974 Miksic 1976 Ozdikmen 2009 Ozdikmen 2009 Islas 1958 Ohaus 1909 Lacroix 1999 Balthasar 1967 Hope 1837 Solier 1851 Coca Abia amp Robbins 2006 Sharp 1875 Ratcliffe amp Cave 2010 Woolley 2016 Burmeister 1842 Fairmaire 1884 Krikken 1976 Guerin Meneville 1840 Dellacasa amp Gordon amp Dellacasa 1998 Westwood 1842 Brenske 1898 Moser 1920 Baraud 1987 Ceraspis Le Peletier amp Serville 1828 Dombrow 2002 Kolbe 1899 Arrow 1948 Ohaus 1911 Arrow 1908 Quedenfeldt 1884 Gestro 1893 Kocak amp Kemal 2008 Ohaus 1912 Cetonia Fabricius 1775 Moser 1917 Heller 1921 Schaum 1862 Westwood 1847 Medvedev 1949 Moser 1914 Arrow 1937 Burmeister 1844 Ohaus 1905 Burmeister 1846 Heller 1903 Dejean 1833 Heller 1902 Burmeister 1844 Chalcosoma Hope 1837 Burmeister 1842 Miksic 1970 Casey 1915 Burmeister 1842 Brenske 1898 Moser 1919 Blanchard 1850 Brenske 1899 Kolbe 1904 Dejean 1821 Le Peletier amp Serville 1828 Saylor 1937 Britton 1990 MacLeay 1864 Britton 1957 Westwood 1842 Lansberge 1875 Hope 1837 Britton 1957 Hope 1840 Vardal amp Forshage 2010 Moron amp Nogueira 1998 Lacroix 1993 Croissandeau 1893 Legrand 2004 Smith amp Mondaca 2016 Sharp 1877 Burmeister 1842 Moser 1924 Chilothorax Motschulsky 1859 Reitter 1905 Kraatz 1891 Soula 2008 Bates 1889 MacLeay 1819 Tarasov 2008 Blanchard 1850 Fairmaire 1905 Bates 1889 Kraatz 1880 Kirby 1828 Burmeister 1844 Burmeister 1844 Kraatz 1880 Kraatz 1880 Westwood 1842 Kolbe 1897 Kirby 1828 Le Peletier amp Serville 1828 Brenske 1897 Burmeister 1847 Soula 2005 Schmidt 1914 Latreille 1829 Schein 1956 Burmeister 1840 Laporte 1832 Cambefort 1984 Burmeister 1842 Lea 1914 Burmeister 1842 Motschulsky 1857 Schoch 1898 Ohaus 1941 Burmeister 1842 Erichson 1847 Lacroix 1999 Fairmaire 1904 Krikken 2009 Li amp Yang 1999 Frey 1970 Arrow 1908 Semenov 1903 Schmidt 1914 Kirby 1827 Kolbe 1910 Motschulsky 1868 Decelle 1979 Paulian 1991 Lacroix 1996 Kolbe 1894 Burgeon 1946 Peringuey 1902 Kolbe 1895 Burmeister amp Schaum 1841 Moser 1913 Hope 1837 Schmidt 1914 Schaum 1841 Horn 1876 Prell 1933 Hope 1831 Ohaus 1910 Ratcliffe amp Hardy 2005 Clement 1958 Paulian 1942 Mulsant 1842 Blanchard 1850 Britton 1987 Erichson 1843 Blackburn 1898 Blanchard 1850 Brenske 1897 Lacroix 2008 Lacroix 1993 Strand 1928 Lacroix 1997 White 1845 Fuhrmann 2012 Kolbe 1884 Fauvel 1903 Erichson 1847 Brenske 1900 Kolbe 1892 Boucomont 1920 Mulsant 1842 Copris Geoffroy 1762 Reiche 1841 Olsoufieff 1924 Balthasar 1938 Harold 1859 Burmeister 1846 Burmeister 1847 Coptomia Burmeister 1842 Pouillaude 1919 Hope 1833 Hope 1845 Moser 1918 Kolbe 1892 Klug 1845 Prell 1936 Kraatz 1880 Saunders 1852 Kraatz 1898 Kolbe 1897 Given 1952 Schurhoff 1933 Cotalpa Burmeister 1844 Burmeister 1842 Kolbe 1895 Schmidt 1914 Blanchard 1851 Knoch 1801 Brenske 1903 Nikolajev 2004 Nikolajev 1993 Nikolajev 1998 Nikolajev 2007 Nikolajev 1995 Nikolajev 1995 Nikolajev 1998 Bordat 2009 Balthasar 1942 Burmeister 1847 MacLeay 1819 Gordon amp Skelley 2007 Kraatz 1888 Britton 1992 Lea 1916 Krikken 1976 Westwood 1874 MacLeay 1838 Cyclocephala Dejean 1821 Klug 1855 Burmeister 1844 Schein 1958 Ohaus 1923 Kirby 1827 Sharp 1875 Waterhouse 1867 Burmeister 1847 Westwood 1842 Fischer Von Waldheim 1823 Brenske 1900 Lesne 1900 Burmeister 1842 Reitter 1903 Brenske 1897 Fairmaire 1895 Kolbe 1895 Brenske 1900 Arrow 1917 Howden amp Storey 2000 Prokofiev 2013 Dechambre 2001 Bordat amp Howden 1995 MacLeay 1819 Moser 1913 Reitter 1902 Le Peletier amp Serville 1828 Endrodi 1952 Kolbe 1904 Allsopp 2022 Lacroix 1993 Lacroix 2005 Lacroix 1999 Sabatinelli amp Pontuale 1998 Vaz de Mello 2008 Gordon amp Skelley 2007 Erichson 1848 Fairmaire 1900 Pereira 1949 Eschscholtz 1822 Harold 1867 Balthasar 1961 Burmeister 1847 Perty 1830 Dechambre 2006 Lacroix 1997 Schein 1959 Dombrow 2002 Arrow 1941 Moser 1915 Ruter 1964 Ocampo amp Mondaca 2012 Alexis amp Delpont 1999 Thomson 1878 Arrow 1907 Gistel 1857 Brenske 1896 De Wailly 1950 Parry 1849 Brown 1929 Harold 1869 Gordon amp Skelley 2007 Newman 1840 Peringuey 1904 Erichson 1847 Burmeister 1844 Horn 1887 Lansberge 1886 Mulsant 1842 Gestro 1888 Waterhouse 1905 Burmeister 1844 Brenske 1895 Blanchard 1850 Burmeister 1855 Harris 1826 Moser 1924 Le Peletier amp Serville 1828 Gory amp Percheron 1833 Burmeister 1847 Dichotomius Hope 1838 Kraatz 1885 Le Peletier amp Serville 1828 Frolov amp Scholtz 2003 Burmeister 1844 Reitter 1903 Dicronocephalus Hope 1831 Hope 1837 Orbigny 1896 Wood mason 1878 Thomson 1877 Balthasar 1959 Gutierrez 1949 Britton 1990 Lacroix 1998 Reiche 1859 Dilochrosis Thomson 1878 Bates 1888 Bordat 2009 Casey 1889 Lacroix 1989 Lacroix 1989 Felsche 1906 Matthews 1974 Hope 1842 Gerstaecker 1882 Le Peletier amp Serville 1828 Fairmaire 1878 Thomson 1858 Hope 1843 Kolbe 1892 Gory amp Percheron 1833 Kirby 1837 Boheman 1857 Burmeister 1842 Burmeister 1842 Burmeister 1842 Harold 1873 Lacroix 1993 Lacroix 1993 Balthasar 1941 Peringuey 1904 Peringuey 1904 Dombrow 2003 Lacroix 1998 Kolbe 1892 Branco 1989 Burmeister 1842 Blanchard 1845 Brenske 1897 Schmidt 1914 Peringuey 1901 Kirby 1828 Boucomont 1921 Deschodt Davis amp Scholtz 2016 Bordat 2009 Carne 1958 Deschodt amp Scholtz 2008 Burmeister 1842 Dynastes MacLeay 1819 Harold 1869 Blackburn 1898 Dechambre 1979 Kraatz 1880 Erichson 1847 Redtenbacher 1868 Ohaus 1915 Neita amp Ocampo 2012 Burmeister 1842 Gene 1836 Moser 1919 Franz 1955 Peringuey 1907 Schmidt 1914 Lacroix 1998 Brenske 1895 Erichson 1847 De Wailly 1950 Erichson 1847 Gerstaecker 1871 Sharp 1877 Erichson 1847 Endrodi 1971 Dejean 1833 Lansberge 1880 Kolbe 1892 Chalumeau 1981 Scholtz amp Howden 1987 Hope 1837 Dalle Torre 1912 Lacroix 2002 Britton 1990 Lansberge 1874 Lucas 1859 Dechambre 1999 Erichson 1847 Moser 1913 Kolbe 1894 Eocorythoderus Marumaya 2012 Barbero Palestrini amp Roggero 2009 Arrow 1908 Carne 1956 Medvedev 1951 Olsoufieff 1947 Reitter 1903 Blanchard 1851 Dejean 1833 Bates 1904 White 1841 Roggero Barbero amp Palestrini 2015 Reiche 1841 Moser 1917 Dejean 1833 Carne 1957 Fairmaire 1880 Tagawa 1941 Burmeister 1842 Hutchinson amp Allsopp 2021 Lacroix 1994 Mulsant 1851 Ohaus 1910 Ohaus 1910 Kolbe 1914 Kolbe 1914 Burmeister 1844 Lacroix 2006 Burmeister amp Schaum 1840 Burmeister 1847 Brenske 1899 Burmeister 1842 Preiss 1902 Schmidt 1917 Mulsant amp Rey 1870 Zidek amp Pokorny 2011 Mulsant amp Rey 1869 Peringuey 1904 Peringuey 1904 Burmeister 1842 Burmeister amp Schaum 1840 Arrow 1907 Burmeister 1842 Prell 1912 Brulle 1837 Waterhouse 1875 White 1839 Burmeister 1840 Mulsant amp Rey 1869 Bates 1888 Bordat 2003 Dellacasa 1983 Lacroix 1997 Fairmaire 1899 Kolbe 1894 Burgeon 1946 Casey 1915 Miksic 1976 Ohaus 1909 Euoniticellus Janssens 1953 Balthasar 1959 Dellacasa 1983 Dellacasa Gordon amp Dellacasa 2016 Le Peletier amp Serville 1828 Brown 1927 Hinton 1936 Stebnicka amp Howden 1996 Eupatorus Burmeister 1847 Duvivier 1892 Brenske 1897 Burmeister 1842 Mulsant 1842 Burmeister 1842 Semenov 1890 Westwood 1867 Marseul 1867 Blanchard 1851 Blanchard 1850 Burmeister 1842 Lacroix 2006 Brenske 1898 Dalman 1824 Moser 1908 Branco 1989 Thomson 1880 Reitter 1896 Waterhouse 1880 Waterhouse 1882 Martinez 1967 Semenov 1890 Castelnau 1840 Szito 1993 Soula 1998 Endrodi 1960 Keith amp Reichenbach 2018 Soula 2002 Carne 1958 Reitter 1902 Reitter 1903 Motschulsky 1859 Soula 2002 Britton 1979 Soula 2002 Bordat Mencl amp Rakovic 2012 Smith amp Mondaca 2015 Martinez 1982 Lacroix 1997 Paulian 1987 Keith amp Montreuil 2004 Blanchard 1850 Vaz de Mello 2008 Dellacasa Dellacasa amp Gordon 2007 Coca Abia 2003 Gordon amp Skelley 2007 Stebnicka 1999 Bertoloni 1852 Howden 1961 Balthasar 1938 Blackburn 1892 Kolbe 1894 Bordat 2022 Burmeister 1842 Antoine 2005 Peringuey 1904 Ratcliffe Jameson amp Zorn 2018 Peringuey 1907 Janssens 1940 Nomura 1973 Brenske 1897 Kirby 1818 Costa Lima 1940 Vaz de Mello 2008 Westwood 1874 Kirby 1825 Gordon amp Skelley 2007 Gordon amp Pittino 1992 Geotrogus Guerin Meneville 1842 Endrodi 1974 Howden 1988 Howden 1980 Janssens 1937 Balthasar 1933 Matsumoto 2014 Moser 1924 Brenske 1899 Lin 1980 Lacroix 1993 Frey 1974 Blackburn 1898 Schoch 1896 Burmeister 1842 Brenske 1899 Westwood 1838 Brenske 1895 Bates 1889 Miksic 1976 Reiche 1860 Reiche 1856 Kirby 1825 Arrow 1914 Erichson 1847 Casey 1915 Lansberge 1887 Kraatz 1880 Krajcik 2009 Gnorimus Le Peletier amp Serville 1828 Endrodi 1982 Janson 1881 Goliathus Lamarck 1801 Thomson 1878 Golofa Hope 1837 Schmidt 1914 Dellacasa Dellacasa amp Gordon 2012 Dellacasa Dellacasa amp Gordon 2012 Reitter 1892 Stebnicka amp Howden 1996 Burmeister 1844 Kolbe 1910 Arrow 1911 Quedenfeldt 1888 Ratcliffe 2010 Skelley Dellacasa amp Dellacasa 2009 Peringuey 1902 Ocampo amp Mondaca 2012 Lacroix 1997 Kolbe 1895 Hutchinson amp Moeseneder 2013 Ziani 2002 Rakovic 1981 Prokofiev 2016 Mlambo Scholtz amp Deschodt 2019 Lacroix 1993 Brulle 1837 Schaeffer 1927 Peringuey 1904 Masumoto amp Kiuchi 2001 Neervoort Van De Poll 1886 Ratcliffe 2019 Casey 1915 Gymnetis MacLeay 1819 Blanchard 1851 Dejean 1833 Arrow 1910 Gymnopleurus Illiger 1803 Linell 1896 Moser 1914 Brenske 1896 Lansberge 1874 Brenske 1893 Kraatz 1898 Britton 1990 Britton 1987 Howden 1995 Kraatz 1892 Moron amp Nogueira 2007 Frey 1967 Cambefort 1978 Roggero Dierkens Barbero amp Palestrini 2017 Halffter amp Martinez 1977 Hope 1837 Kolbe 1910 Arrow 1908 Semenov 1890 Peringuey 1901 Harold 1861 Bordat 2003 Chalumeau 1981 Gordon amp Skelley 2007 Boucomont 1914 Britton 1957 Harpina Burmeister 1844 Burmeister 1847 Sharp 1877 Kolbe 1894 Thomson 1881 Orbigny 1915 Heliocopris Hope 1837 Baraud amp Nicolas 1966 Krikken 1982 Brenske 1897 Baraud 1986 Miksic 1970 Kraatz 1880 Tangelder amp Krikken 1982 Reitter 1897 Schmidt 1914 Reiche 1860 Miksic 1974 Kraatz 1880 Miksic 1972 Burmeister 1842 Kolbe 1910 Miksic 1963 Mulsant 1842 Brenske 1898 Motschulsky 1857 Seilliere 1910 Antoine 1989 Burmeister 1844 Soula 2002 Burmeister 1842 Peringuey 1901 Albers 1852 Kolbe 1900 Martinez 1948 Bourgoin 1919 Ohaus 1909 Moser 1911 Dupuis amp Dechambre 2008 Burmeister 1847 Kolbe 1900 Gillet 1911 Heteronychus Dejean 1833 Guerin Meneville 1830 Burmeister 1842 Blanchard 1851 Valck Lucassen 1933 Westwood 1842 Moser 1917 Brenske 1899 Schaum 1844 Dupont 1832 Dupuis 2006 Antoine 2002 Moser 1918 Krikken 1978 Fairmaire 1891 Olivier 1789 Fairmaire 1901 Endrodi 1978 Burmeister 1855 Fairmaire 1886 Sabatinelli 1983 Howden amp Endrodi 1978 Nikritin 1977 Peringuey 1904 Martinez amp Pereira 1956 Hope 1838 Reitter 1889 Brenske 1894 Martinez 1949 Brenske 1891 Brenske 1896 Britton 1978 Hope 1837 Blanchard 1851 Brenske 1897 Janssens 1932 Soula 2006 Burmeister 1842 Burmeister 1847 Kolbe 1894 Ohaus 1901 MacLeay 1871 Guerin Meneville 1839 Kolbe 1910 Lacroix 1998 Ohaus 1898 Kolbe 1895 Brenske 1899 Kunckel D Herculais 1887 Hoplia Illiger 1803 Blanchard 1850 Kolbe 1904 Blanchard 1851 Harold 1869 Fairmaire 1904 Nomura 1974 Bates 1904 Laporte 1832 Thomson 1880 Schurhoff 1933 Moser 1918 Prell 1933 Burgeon 1946 Stebnicka 2000 Dellacasa Dellacasa amp Gordon 2015 Endrodi 1974 Fairmaire 1897 Arrow 1911 Hardy 1978 Arnaud 1993 Lacroix 1993 Palestrini amp Giacone 1988 Quedenfeldt 1884 Dejean 1833 Brenske 1898 Peringuey 1901 Peringuey 1904 Nikolajev 1996 Kolbe 1904 Arrow 1899 Ohaus 1910 Reitter 1890 Eschscholz 1830 Arrow 1899 Fairmaire 1878 Kraatz 1880 Blackburn 1895 Balthasar 1938 Britton 1978 Brenske 1897 Howden 1968 LeConte 1861 Burmeister 1840 Martinez 1953 Rossner 2018 Gory amp Percheron 1833 Britton 1986 Peringuey 1904 Peringuey 1904 Britton 1987 Arrow 1916 Arrow 1916 Arrow 1932 Arrow 1910 Peringuey 1902 Heller 1916 Stebnicka 1997 Montreuil amp Lacroix 2020 Peringuey 1902 Le Peletier amp Serville 1828 Thomson 1857 Dechambre 1979 Mencl Kral amp Rakovic 2019 Krikken 2009 Krikken 2009 Ricchiardi 2012 Endrodi 1976 Fairmaire 1893 Smith amp Mondaca 2015 Nikolajev 2006 Ratcliffe 2003 Delgado amp Moron 1991 Gordon amp Skelley 2007 Pacheco amp Vaz de Mello 2019 Gestro 1874 Kraatz 1892 Burmeister 1842 Krikken 1978 Kraatz 1880 Kolbe 1897 Pereira amp Martinez 1960 Mannerheim 1828 Burmeister 1855 Fairmaire 1889 Roth 1851 Thomson 1880 Balthasar 1967 Dellacasa Gordon amp Dellacasa 2003 Schurhoff 1937 Paulian 1976 Cambefort 1975 Dierkens Moretto amp Genier 2017 Lacroix 1993 Saunders 1839 Dalla Torre 1913 Nikolajev 2005 Lacroix 2005 Frey 1965 Peringuey 1902 Minkina 2017 Perissinotto 2020 Lacroix 2011 Lacroix 2009 Delpont 1996 Janssens 1940 Peringuey 1902 Nagai 1984 Allsopp 2022 Peringuey 1902 Jacobson 1906 Lacroix 1993 Kim 1996 Peringuey 1902 Schmidt 1914 Peringuey 1904 Branco 1991 Nikolajev amp Kabakov 1977 Peringuey 1902 Lacroix 1989 Roggero Barbero amp Palestrini 2017 Mulsant amp Rey 1869 Sharp 1873 Lacroix 1997 Erichson 1847 Reitter 1889 Keith 2000 Keith 2007 Hoffmannsegg 1817 Krikken 1980 Brenske 1897 Ritsema 1888 Keith 2007 Balthasar 1963 Semenov Tian Shanskii amp Medvedev 1936 Blanchard 1851 Erichson 1847 Brenske 1897 Reitter 1899 Reitter 1913 Moser 1921 Krikken 2009 Lacroix 1993 Kolbe 1907 Kolbe 1894 Gordon amp Skelley 2007 Rakovic 1981 MacLeay 1863 Britton 1986 Fabrizi Eberle amp Ahrens 2019 Vaz de Mello 2008 Balthasar 1966 Lepidiota Kirby 1828 Kolbe 1894 Nikolajev 1979 Kolbe 1894 Moser 1914 Le Peletier amp Serville 1828 Le Peletier amp Serville 1828 Schein 1959 Moser 1914 Baraud amp Branco 1991 Dombrow 2001 Saylor 1935 Howden 2003 Peringuey 1904 Burmeister 1842 Dejean 1833 Brenske 1892 MacLeay 1819 Erichson 1847 Reitter 1892 Erichson 1835 Gutierrez 1946 Martinez 1953 Prell 1933 Burmeister 1847 Mulsant amp Rey 1869 Lacroix 1999 Rakovic amp Mencl 2012 Petrovitz 1961 Guerin Meneville 1830 Pittino 2006 Hutchinson amp Allsopp 2022 Burmeister 1842 Motschulsky 1859 Kolbe 1892 Guerin Meneville 1830 Arrow 1917 Schaum 1844 Bates 1889 Blanchard 1851 Nikolajev 1992 Nikolajev 1992 Waterhouse 1891 Montreuil Genier amp Nel 2010 Ohaus 1917 Schmidt 1914 Kolbe 1914 Kolbe 1897 Martinez 1951 Stebnicka 1999 Gory amp Percheron 1833 Kolbe 1895 Burmeister 1847 Endrodi 1980 Frey 1974 Janssens 1938 Westwood 1842 Perissinotto Clennell amp Beinhundner 2019 Reitter 1892 Mulsant amp Rey 1869 Peringuey 1901 Frey 1969 Miksic 1968 Olsoufieff 1940 Martinez 1972 Balthasar 1964 Westwood 1875 Ozdikmen 2009 Gordon amp Skelley 2007 Breme 1844 Kraatz 1880 Lacroix 1993 Britton 1990 MacLeay 1819 Schurhoff 1935 Ohaus 1910 Macrodactylus Latreille 1825 Westwood 1842 Westwood 1874 Hoffmannsegg 1817 Krikken 1977 Brenske 1898 Lacroix 1997 Moron 1983 Guerin Meneville 1844 Ohaus 1905 Bordat Dellacasa amp Dellacasa 2000 Peringuey 1908 Strand 1934 Gordon amp Skelley 2007 Lacroix 1998 Paulian 1992 Ratcliffe amp Rome 2019 Chalumeau amp Gruner 1976 Lea 1919 MacLeay 1819 Montreuil amp Keith 2022 Lacroix 1993 Peringuey 1908 Lacroix 1993 Mulsant 1871 Martinez 1961 Heller 1891 Arrow 1941 Lacroix 2013 Blanchard 1850 Paulian 1974 Moser 1919 Moser 1919 Castelnau 1840 Moser 1924 Kirby 1827 Thomson 1880 Coquerel 1866 Chalumeau amp Ozdikmen 2006 Vaz de Mello 2008 Arrow 1919 Hirasawa 1992 Gunter amp Weir 2017 Dombrow 2002 Lansberge 1882 Alexis amp Delpont 2001 Lacroix 1993 Moron amp Nogueira 1998 Antoine 1997 Bates 1904 Mulsant 1870 Hope 1837 Keith 2001 Hope 1837 Dechambre 1976 Britton 1987 Janssens 1937 Reitter 1903 Schurhoff 1933 Britton 1978 Megasoma Kirby 1825 Kakizoe Maruyama amp Masumoto 2019 Landin 1974 Reitter 1892 Waterhouse 1891 Eschscholtz 1822 Balthasar 1939 Burmeister 1855 Lacroix 1997 Kraatz 1900 Fairmaire 1881 Brenske 1899 Melanocanthon Halffter 1958 Lin 1980 Melinopterus Mulsant 1842 Wollaston 1869 Melolontha Fabricius 1775 Heer 1847 De Wailly 1950 Reitter 1901 Harold 1868 Arrow 1941 Castelnau 1840 Brenske 1897 Krikken 1982 Lacroix 1993 Gordon amp Skelley 2007 Thomson 1880 Ohaus 1905 Motschulsky 1863 Brenske 1897 Moron 1987 Kolbe 1914 Rakovic amp Kral 1997 Ponomarenko 1977 Waterhouse 1878 Paulian 1939 Machatschke 1957 Kraatz 1880 Valck Lucassen 1933 Arrow 1911 Bates 1889 Kolbe 1910 Lacroix 2008 Westwood 1842 Britton 1978 Frolov amp Akhmetova 2017 Ruter 1969 Lacroix 1997 Brown 1931 Arrow 1911 Pouillaude 1917 Moser 1910 Blanchard 1851 Britton 1957 Balthasar 1958 Jakl 2009 Burmeister 1844 Kraatz 1885 Petrovitz 1967 Gutierrez 1951 Blanchard 1842 Baraud 1991 Kolbe 1910 Kraatz 1890 Peringuey 1902 Burmeister 1844 Kraatz 1880 Lea 1920 Bates 1904 Arrow 1908 Brenske 1894 Nikolajev 1979 Pittino 2006 Landin 1974 Burmeister 1855 Brenske 1900 Kraatz 1883 Ohaus 1911 Montreuil amp Keith 2022 Peringuey 1901 Arrow 1901 Nikolajev 2007 Kirby 1823 Casey 1915 Martinez 1964 Balthasar 1963 Arrow 1944 Soula 1998 Arrow 1902 Lacroix 1998 Wickham 1914 Lacordaire 1855 Ruter 1953 Reitter 1902 Brenske 1903 Thomson 1859 Burmeister 1844 Ritsema 1888 MacLeay 1821 Fairmaire amp Germain 1860 Kraatz 1897 Antoine 1989 Nikolajev 2004 Le Peletier amp Serville 1828 Lansberge 1874 Erichson 1847 Howden amp Storey 2000 Gunter amp Weir 2017 Roggero Dierkens Barbero amp Palestrini 2017 Kraatz 1880 Janson 1905 Ramirez Ponce 2015 Grossi amp Vaz de Mello 2015 Ahrens Lukic amp Liu 2023 Ruter 1965 Semenov amp Medvedev 1927 Huang amp Chen 2016 Nomura amp Nakane 1951 Soula 2002 Lacroix 1993 Vauloger 1898 Broun 1904 Neervoort Van De Poll 1886 Berg 1881 Martinez 1975 Ruter 1964 Bourgoin 1920 Burmeister 1840 Balthasar 1955 Martinez 1952 Wasmann 1900 Petrovitz 1962 Burmeister 1842 Perissinotto 2020 Hirasawa 1992 Scholtz amp Howden 1987 Deschodt amp Davis 2017 Peringuey 1902 Burmeister 1844 Kolbe 1894 Britton 1978 Westwood 1842 Howden 2003 Stebnicka 1999 Dupont 1835 Moeseneder amp Hutchinson 2016 Moron amp Nogueira 1998 Koshantschikov 1912 Deschodt amp Scholtz 2008 Brenske 1900 Saylor 1938 Arrow 1906 Arrow 1917 Bordat 1990 Burgeon 1945 Perissinotto 2017 Neervoort Van De Poll 1886 Lacroix 1997 Sharp 1878 Landin 1974 Ratcliffe amp Mico 2001 Carne 1985 Carne 1957 Dellacasa 1984 Clement 1986 Martinez 1953 Paulian amp Villiers 1941 Blackburn 1890 Heller 1896 Baraud 1965 Yamaya 2010 Minkina 2023 Alexis amp Delpont 1998 Carne 1957 Peringuey 1901 Lacroix 2015 Lucas 1870 Allsopp 1983 Blackburn 1898 Rakovic 1986 Gordon amp Pittino 1992 Thomson 1878 Arrow 1908 Branco 1990 Schoch 1897 Brenske 1894 Miyake amp Yamaya 1994 Dellacasa Dellacasa amp Skelley 2010 Rakovic amp Kral 1997 Dellacasa Gordon amp Dellacasa 2004 Miyake 1985 Frey 1965 Sharp 1873 Erichson 1842 Prokofiev 2016 Blackburn 1898 Scott 1912 Vinson 1946 Martinez amp Pereira 1958 Martinez 1952 Smith amp Mondaca 2015 Brenske 1900 Kolbe 1908 Hollande amp Therond 1998 Mulsant amp Rey 1869 Dombrow 2002 Matsumoto 2016 Paulian 1970 Nimbus Mulsant amp Rey 1869 Kraatz 1883 Kraatz 1880 Nakane 1959 Ochi amp Kawahara 2001 Nomura 1973 Sawada 1941 Branco 2010 Britton 1957 Allsopp amp Hutchinson 2020 Dellacasa amp Dellacasa 2005 Mulsant amp Rey 1869 Petrovitz 1962 Blackburn 1898 Heller 1896 Quedenfeldt 1884 Sharp 1875 Holm amp Perissinotto 2004 Prokofiev 2016 Vaz de Mello 2008 Westwood 1879 Bordat 1988 Lacroix 1998 Vaz de mello 2003 Moeseneder amp Hutchinson 2012 Brenske 1892 Harold 1877 Dellacasa Kral Dellacasa amp Bordat 2014 Schmidt 1917 Boheman 1857 Koshantschikov 1916 Fairmaire 1897 Gordon amp Pittino 1992 Burmeister 1842 MacLeay 1871 Kral Rakovic amp Mencl 2015 White 1846 Waterhouse 1875 Burmeister 1855 Kolbe 1897 Kraatz 1894 Schonherr 1817 Schein 1958 Burmeister 1844 LeConte 1856 Kolbe 1891 Dejean 1821 Onitis Fabricius 1798 Erichson 1847 Vaz de Mello 2008 Frolov amp Vaz de Mello 2015 Erichson 1847 Reiche 1860 Onthophagus Latreille 1802 Arrow 1914 Moser 1916 Frey 1972 Boucomont 1912 Peringuey 1904 Solier 1851 Britton 1987 MacLeay 1819 MacLeay 1838 Britton 1957 Kral Rakovic amp Mencl 2016 Burmeister 1842 Lacroix 1998 Kolbe 1897 Kolbe 1904 Fairmaire 1878 Endrodi 1983 Krikken 1982 Schmidt 1914 Reitter 1900 Mulsant amp Rey 1869 MacLeay 1819 Ohaus 1912 Burmeister 1847 Carne 1957 Brenske 1900 Reitter 1895 Bates 1870 Oryctes Hellwig 1798 Ratcliffe Smith amp Erwin 2005 Endrodi 1978 Boisduval 1835 Guerin Meneville 1830 Kolbe 1910 Gordon amp Skelley 2007 Branco amp Baraud 1988 Le Peletier amp Serville 1828 Brenske 1896 Mulsant 1842 D Andretta amp Martinez 1957 Scholtz amp Howden 1987 Peringuey 1902 Costa Cherman amp Iannuzzi 2020 Frey 1968 Arrow 1917 Dellacasa amp Stebnicka 2001 Quedenfeldt 1888 Solsky 1876 Arrow 1908 Dellacasa Dellacasa amp Gordon 2016 Dejean 1833 Pouillaude 1920 Brenske 1899 Oxysternon Castelnau 1840 Mulsant 1842 Skelley 2007 Soula 2006 Brenske 1894 Pachnoda Burmeister 1842 Alexis amp Delpont 2002 Blanchard 1851 Brenske 1895 Brenske 1903 Le Peletier amp Serville 1828 Erichson 1847 Laporte 1832 Reitter 1902 Moser 1920 Germar 1848 Griffith amp Pidgeon 1831 Arrow 1908 Fairmaire 1884 Dejean 1821 Ohaus 1912 Brenske 1897 MacLeay 1821 Blanchard 1851 De Palma amp Malec 2020 Hope 1841 Curtis 1845 Lacroix 1999 Kolbe 1910 Reiche 1871 Thomson 1880 Poinar 2011 Stebnicka 1985 Lewis 1895 Kraatz 1880 Reitter 1902 Burmeister 1847 Burmeister 1842 Papuana Arrow 1911 Hollande amp Therond 1998 Ohaus 1915 Peringuey 1904 Balthasar 1938 Dombrow 2002 Burmeister 1842 Soula 2002 Harold 1873 Bates 1888 Waterhouse 1875 Balthasar 1963 Wasmann 1918 Ohaus 1915 Balthasar 1932 Howden amp Cook 2002 Howden 1971 Landin 1959 Balthasar 1937 Bordat 2022 Machatschke 1957 Soula 1998 Vulcano Pereira amp Martinez 1966 Becker 1910 Shipp 1897 Moser 1924 Moron 1983 Lacroix amp Coache 2017 Roggero Barbero amp Palestrini 2015 Antoine 1989 Ocampo amp Colby 2009 Ohaus 1915 Nomura 1974 Lacroix 1998 Seidlitz 1888 Lacroix 1997 Antoine 1988 Schmidt 1914 Cockerell 1905 Casey 1915 Martinez 1961 Balthasar 1969 Endrodi 1978 Harold 1877 Arrow 1910 Miksic 1985 Chalumeau 1992 Hauser 1904 Moser 1907 Verdu Stebnicka amp Galante 2006 Soula 2002 Prell 1934 Kraatz 1899 Blackburn 1911 Paulian 1991 Balthasar 1955 Paulian 1932 Ohaus 1904 Britton 1990 Reitter 1896 Westwood 1842 Antoine 2005 Pacheco Monne Vaz de Mello amp Ahrens 2022 Balthasar 1961 Ohaus 1915 Shipp 1895 Ohaus 1915 Bordat 2013 Janson 1881 Baraud 1962 Lacroix Coache amp Filippi 2022 Holm 1988 Gordon amp Skelley 2007 Marais amp Holm 1989 Kraatz 1893 Moser 1918 Prell 1934 Kraatz 1880 Ohaus 1915 Ohaus 1915 Bates 1888 Schaufuss 1890 Balthasar 1963 Paulian 1984 Britton 1990 Fairmaire 1903 Dechambre 1994 Brenske 1899 Deschodt amp Scholtz 2008 Peringuey 1904 Balthasar 1945 Soula 2008 Balthasar 1938 Martinez amp D andretta 1956 Scholtz amp Howden 1987 Ballion 1871 Arrow 1913 Laporte 1832 Harold 1868 Kraatz 1880 Matsumoto 2016 Erichson 1847 Pelidnota MacLeay 1819 Cockerell 1920 Burmeister 1847 Paulian 1985 Moser 1926 Ohaus 1919 De Wailly 1950 Brenske 1896 Brenske 1891 Kraatz 1892 Endrodi 1968 Hope 1837 Peringuey 1901 Krikken 1983 Westwood 1847 Vaz de Mello 2008 Peringuey 1904 Brenske 1896 Burmeister 1847 Kraatz 1880 Lacroix 2006 Kolbe 1910 Brenske 1898 Waterhouse 1875 Burmeister 1844 Lacroix 1997 Mondaca amp Valencia 2016 Blackburn 1898 Miksic 1965 Rakovic 1979 Reitter 1903 Reitter 1892 Waterhouse 1841 Paulian 1985 Hope 1841 Hope 1833 Motschulsky 1859 Burmeister 1844 Reitter 1889 Quedenfeldt 1884 Erichson 1848 MacLeay 1819 Kraatz 1894 Blanchard 1851 Reitter 1892 Brenske 1897 Heller 1900 Dechambre 2008 Latreille 1807 Zidek 2021 MacLeay 1838 Dejean 1833 Brethes 1919 Brenske 1897 LeConte 1856 Bordat 2009 Kolbe 1894 Moser 1919 Kraatz 1880 Britton 1987 Brenske 1897 Schaum 1844 Schoch 1898 Broun 1886 Blackburn 1890 Moser 1918 Blanchard 1846 Waterhouse 1876 Eschscholtz 1830 Stephens 1830 Phyllophaga Harris 1826 Brenske 1899 Blackburn 1898 Fischer Von Waldheim 1823 Brenske 1900 Blanchard 1851 Soula 2002 Bourgoin 1921 Kraatz 1888 Burmeister 1842 Moser 1924 Erichson 1842 Branco 1991 Harold 1875 Ratcliffe amp Giraldo 2014 Rakovic amp Kral 1997 Peringuey 1900 Brenske 1899 Mulsant 1842 Dellacasa amp Dellacasa 2005 Dellacasa amp Dellacasa 2005 Mulsant amp Rey 1869 Chapin 1932 Burmeister 1844 Dejean 1833 Schmidt 1917 MacLeay 1819 Krikken 1978 Westwood 1854 Janssens 1942 Ohaus 1910 Bates 1888 Westwood 1874 Mulsant 1842 Le Peletier amp Serville 1828 Horn 1867 Wallace 1867 Peringuey 1904 Erichson 1847 Lacroix amp Montreuil 2019 Lacroix 2006 Jameson 1997 Moron 1983 Schmidt 1914 Lansberge 1875 Kraatz 1892 Mulsant 1842 Fairmaire 1896 Ruter 1978 Brenske 1899 Brenske 1899 Burmeister 1847 Burmeister 1847 Harold 1869 Moser 1915 Moser 1917 Howden 1997 Paulian 1942 Schmidt 1914 Arrow 1910 Poecilopharis Kraatz 1880 Kraatz 1880 Arrow 1910 Burmeister 1842 Waterhouse 1875 De Wailly 1950 Polyphylla Harris 1841 Blanchard 1851 Janson 1888 Kraatz 1882 Popillia Dejean 1821 Burmeister 1842 Arrow 1911 Krikken 1984 Skelley 2021 Jakl 2018 Harold 1869 Lansberge 1883 Reitter 1903 Blanchard 1851 Deyrolle 1864 Britton 1988 Erichson 1847 Harold 1861 Lopez Garcia amp Deloya 2022 Broun 1904 Brenske 1893 Sigwalt 1987 Bai Ren amp Yang 2011 Newman 1836 Kolbe 1905 Protaetia Burmeister 1842 Miksic 1968 Britton 1987 Brenske 1896 Krikken 1976 Krikken 1978 Decelle 1979 Nikolajev 2007 Nikolajev 2000 Nikolajev Wang amp Zhang 2014 Endrodi 1976 Fallen 1807 Thomson 1859 Stebnicka 1994 Brenske 1899 Peringuey 1904 Reitter 1892 Semenov 1890 Heller 1923 Schmidt 1914 Antoine 1989 Bourgoin 1921 Brown 1927 Fairmaire 1901 Lacroix 1991 Kraatz 1880 Hollande amp Therond 1998 Orbigny 1896 Schein 1959 Blackburn 1911 Paulian 1984 Kraatz 1880 Frey 1971 Bordat 1994 Dellacasa amp Dellacasa 1997 Soula 1998 Bordat 2016 Keith 2000 Bates 1887 Blackburn 1890 Ritsema 1882 Miksic 1985 Ferreira 1977 Ohaus 1905 Kraatz 1882 Neervoort Van De Poll 1886 Dellacasa Dellacasa amp Gordon 2013 Moser 1924 Ferreira 1960 Hardy 1971 Frolov amp Montreuil 2019 Miksic 1974 Gutierrez 1950 Nonfried 1894 Soula 1998 Bordat 1993 Ohaus 1910 Dellacasa Gordon amp Dellacasa 2007 Dellacasa amp Dellacasa 2000 Blackburn 1892 Arrow 1908 Soula 1998 Moser 1919 Ohaus 1903 Nomura 1974 Ohaus 1912 Lacroix 1998 Pittino 1984 Minkina 2023 Paulian 1984 Baraud 1985 Balthasar 1930 Petrovitz 1969 Lacroix 1997 Katovich 2011 Felsche 1904 Keith amp Reichenbach 2018 Bordat 1990 Duvivier 1892 Moser 1918 Dellacasa 1990 Soula 2002 Benderitter 1913 Sharp 1873 Balthasar 1941 Krikken 1976 Ohaus 1904 Keith 2006 Guerin Meneville 1830 Kobayashi 1980 Ricchiardi 2015 Bordat amp Howden 1995 Dalle Torre 1912 Peringuey 1901 Kraatz 1882 Dellacasa Dellacasa amp Gordon 2015 Dellacasa 1986 Soula 2002 Keith 2008 Miksic 1967 Jacquelin Du Val 1859 Keith 2007 Quedenfeldt 1884 Schein 1959 Petrovitz 1962 Peringuey 1904 Burmeister 1842 Broun 1895 Burmeister 1855 Brenske 1892 Bates 1888 Murayama 2010 Kraatz 1883 Redtenbacher 1868 Mulsant amp Rey 1869 Ohaus 1910 Ocampo amp Smith 2006 Muramoto 2006 Smith 2008 Arrow 1931 Thomson 1859 Burmeister 1842 Kolbe 1884 Boisduval 1835 Pyrrhopoda Kraatz 1880 Kral 1997 Peringuey 1902 Thomson 1877 Pittino 2006 Kolbe 1894 Lacroix 1991 Saylor 1947 Schein 1958 Fairmaire 1860 Reineria Miksic 1968 Lacroix 1993 Balthasar 1951 Frolov amp Montreuil 2009 MacLeay 1819 Burmeister 1855 Burmeister 1842 Krikken 1981 Lacroix 1997 Arrow 1930 Kraatz 1900 Burmeister 1842 Ohaus 1912 Klug 1855 Ruter 1965 Perty 1833 Maruyama 2009 Burmeister 1842 Burmeister 1844 Ohaus 1909 Peringuey 1901 Westwood 1841 Kolbe 1894 Rhizotrogus Latreille 1825 Adam 1994 Hope 1837 Erichson 1847 Ohaus 1913 Lacroix 2008 Fairmaire 1883 Burgeon 1942 Agassiz 1846 Bedel 1911 Reitter 1892 Clouet 1900 Mulsant 1842 Moser 1921 Moser 1924 Burmeister 1842 Hutchinson amp Moeseneder 2022 Lacroix 1997 Lacroix 1996 De Wailly 1950 Matsumoto 2016 Gordon amp Skelley 2007 Machatschke 1957 Latreille 1802 Waterhouse 1874 Bates 1888 Arrow 1908 Krikken 1980 Montreuil amp Lacroix 2020 Lacroix 1998 Moron amp Grossi 2015 Dellacasa amp Dellacasa 2000 Brookes 1944 Sharp 1873 Stebnicka amp Galante 2007 Balthasar 1967 Redtenbacher 1857 Stebnicka 1993 Blackburn 1904 Rakovic amp Mencl 2011 Erichson 1847 Blackburn 1907 Waterhouse 1878 Paulian 1934 Frey 1975 Gordon amp Skelley 2007 Burmeister 1847 Evans 1987 Quedenfeldt 1884 Ohaus 1912 Schaum 1841 Peringuey 1901 Balthasar 1965 Scarabaeus Linnaeus 1758 Erichson 1847 Erichson 1835 Genier amp Kohlmann 2003 Westwood 1837 Dombrow 2001 Burmeister 1844 Gordon amp Skelley 2007 Ruter 1957 Gautier Des Cottes 1872 Arrow 1917 Blanchard 1850 Fischer Von Waldheim 1823 Dejean 1833 Kirby 1825 Berg 1898 Erichson 1842 Britton 1990 Blackburn 1892 Martinez 1948 Martinez 1953 Broun 1886 Kraatz 1880 Peringuey 1904 Castelnau 1840 Brenske 1897 Fairmaire 1891 Stebnicka 1999 Hope 1847 De Wailly 1950 Serica MacLeay 1819 Motschulsky 1860 Boisduval 1835 Guerin Meneville 1839 Sharp 1882 Kabakov 1996 Miyake 2002 Gordon amp Skelley 2007 Stebnicka 1994 Masumoto 1991 Keith 2005 Reitter 1896 Mulsant amp Rey 1869 Landin 1949 Roets amp Oberlander 2010 Vaz de Mello 2008 Harold 1871 Kraatz 1886 Cervenka 1994 Boucomont 1928 Blanchard 1851 Dellacasa 1986 Simonis 1985 Dechambre 1998 Sisyphus Latreille 1807 Fairmaire 1894 Dellacasa Dellacasa Gordon amp Stebnicka 2011 Westwood 1847 Kral Sehnal amp Bezdek 2012 Lansberge 1882 Brenske 1899 Fairmaire 1887 Soula 2006 Kolbe 1894 Castelnau 1840 Arrow 1902 Lansberge 1886 Kakizoe Jiang amp Wang 2021 Jameson 1997 Burmeister 1855 Brenske 1900 Brenske 1899 Kolbe 1904 Kraatz 1890 Sharp 1877 Westwood 1848 Kraatz 1892 Ohaus 1910 Machatschke 1970 Burmeister 1847 Burmeister 1855 Burmeister 1842 Skelley 2007 Burmeister amp Schaum 1840 Britton 1990 Burmeister 1847 Fairmaire 1889 Arrow 1943 Moser 1913 Brenske 1900 Nomura 1977 Karsch 1881 Burmeister 1842 Schmidt 1914 Brenske 1896 Stephanorrhina Burmeister 1842 Arrow 1905 Hope 1837 Bainbridge 1840 Valck Lucassen 1933 Dombrow 2001 Kolbe 1914 Branco 1989 Harold 1871 Branco 1989 Kolbe 1897 Hasenpusch amp Moeseneder 2010 Fairmaire 1901 Strategus Hope 1837 Lansberge 1874 Burmeister 1844 Gordon amp Skelley 2007 Gory amp Percheron 1833 Ohaus 1912 Mulsant amp Rey 1869 Reitter 1894 Branco 1992 Olsoufieff 1924 Peringuey 1904 Mencl 2011 Martinez 1955 Paulian 1954 Fairmaire 1896 Boheman 1857 Halffter amp Martinez 1977 Erichson 1835 Schmidt 1914 Fairmaire 1899 Bates 1868 Britton 1956 Kolbe 1897 Lacroix 1993 Brenske 1898 Kolbe 1897 Peringuey 1901 Peringuey 1901 Janssens 1942 Sharp 1877 Kraatz 1895 Burmeister 1842 Valck Lucassens 1939 Kobayashi 1990 Yamaya 2001 Brenske 1899 Stebnicka 2006 Petrovitz 1971 Menetries 1832 Lacroix 2008 Lacroix 2002 Scholtz amp Howden 1987 Quedenfeldt 1888 Thomson 1880 Moser 1924 Miyake 1993 Burmeister 1842 Endrodi 1975 Sharp 1873 Burmeister 1844 Krikken 1975 Erichson 1842 Britton 1990 Westwood 1841 Hope 1837 Burmeister 1842 Brenske 1900 Brenske 1897 Lacroix 1989 Krikken 1970 Mateu 1966 Nakane 1961 Wasmann 1894 Britton 1957 Wasmann 1918 Reichensperger 1915 Krikken 2018 Rakovic 1981 Hope 1837 Vaz de Mello amp Halffter 2006 Matsumoto 2021 Schmidt 1914 Blanchard 1845 Kraatz 1896 Duarte amp Grossi 2022 Blanchard 1842 Ahrens 2004 Mulsant 1842 Peringuey 1902 Thomson 1880 Huchet 2000 Soula 2006 Semenov amp Reichardt 1925 Prokofiev 2015 Decelle 1968 Burmeister 1847 Brenske 1897 LeConte 1856 Blackburn 1904 Reitter 1898 Burmeister 1844 Ohaus 1915 Prokofiev 2014 Burmeister 1842 Roggero Moretto Barbero amp Palestrini 2019 Krikken 2009 Miyake 1994 Krikken 1978 Peringuey 1901 Soula 2002 Peringuey 1904 Westwood 1841 Reitter 1902 Erichson 1847 Orbigny 1904 Prokofiev 2021 Arrow 1907 Pavicevic 1985 Dombrow 2002 Fairmaire 1878 Brenske 1899 Tragiscus Klug 1855 Faldermann 1835 Kolbe 1914 Dupuis amp Perrin 2022 Reitter 1903 Arrow 1917 Schmidt 1914 Paulian 1942 Schmidt 1914 Kraatz 1880 Erichson 1847 Vaz de Mello 2008 Harold 1868 Waterhouse 1875 Minkina 2018 Burmeister 1855 Petrovitz 1963 Clouet 1901 Trichiotinus Casey 1915 Trichius Fabricius 1775 Moser 1916 Burmeister 1847 Blanchard 1851 Nomura 1952 Nomura 1974 Legrand 2001 Balthasar 1945 Bedel 1911 Bordat 2022 Burmeister 1842 Ohaus 1897 Moser 1917 Burmeister 1842 Brenske 1896 Kraatz 1894 Burmeister amp Schaum 1840 Pouillaude 1913 Hope 1831 Petrovitz 1963 Brenske 1893 Brenske 1894 Reitter 1919 Moser 1921 Westwood 1845 Saylor 1942 Ohaus 1904 Burmeister 1847 Moser 1922 Krikken 1987 Burmeister 1847 Balthasar 1932 Ohaus 1917 Reitter 1896 Brenske 1900 Laporte 1832 Boheman 1857 Janssens 1937 Mulsant 1842 Blanchard 1851 Gerstaecker 1867 Minck 1920 Lacroix 1997 Peringuey 1904 Semenov 1905 Ochi amp Kon 2017 Saylor 1945 Burmeister 1842 Josso amp Prevost 2008 Deschodt Sole amp Scholtz 2020 Westwood 1842 De Wailly 1950 Ruter 1973 Fairmaire 1899 Valgus Scriba 1790 Schein 1958 Lacroix 1993 Ohaus 1915 Deschodt Davis amp Scholtz 2011 Lacroix 1993 Ruter 1964 Jameson 1990 Dellacasa Dellacasa amp Bordat 2002 Volinus Mulsant amp Rey 1869 Itoh 1993 Allsopp Jakl amp Rey 2023 Cambefort 1977 Allsopp 1988 Smith amp Evans 2005 Britton 1995 Britton 1988 Dechambre 1999 Reitter 1902 Kraatz 1881 Arrow 1915 Martinez 1952 Arrow 1920 Soula 2002 Breme 1844 Villatoro amp Jameson 2001 Hinton 1934 Bates 1889 Bates 1904 Kolbe 1896 Arrow 1910 Ahrens 2005 Moser 1917 Britton 1978 Saylor 1937 Harold 1869 Burmeister 1842 Holm 1992 Boisduval 1835 Hope 1837 Prell 1934 Xylotrupes Hope 1837 Britton 1990 Britton 1987 Britton 1987 Britton 1987 Moron amp Nogueira 1998 Nomura 1952 Nikolajev 2015 Bordat 1988 Ochi amp Kon 1996 Saylor 1945 Fuhrmann 2021 Fairmaire 1894 Blackburn 1894 Lacroix 1997 Lacroix 2006 Arrow 1932Les scarabees dans la cultureCollection d amulettes en forme de scarabees Le Scarabee sacre Scarabaeus sacer est un animal venere dans l Egypte antique Symbole du soleil et de Re il etait l un des douze animaux sacres associe aux douze heures du jour et de la nuit ref necessaire Le Scarabee rhinoceros siamois ou scarabee de combat Xylotrupes gideon est capture dans la nature puis dresse pour des duels qui font l objet de paris dans le nord de la Thailande Ces duels de scarabees sont couramment organises Ces combats d insectes du type xylotrupes appeles Kwang localement sont a bien des egards surprenants notamment du fait qu il est sans doute plus difficile d attribuer a un scarabee qu a un autre animal des caracteristiques anthropomorphiques et qu il est plus complique encore de deviner ce qu il percoit Le duel de scarabees met a l epreuve une forme de zoomanite trouble fondee sur une forme d attachement inedite entre l homme et l animal Notes et referencesMuseum de l universite du Nebraska Michael Chinery Insectes de France et d Europe occidentale Flammarion 2012 1re ed 1986 p 264 Anatole Bailly 2020 Hugo Chavez Gerard Greco Andre Charbonnet Mark De Wilde Bernard Marechal amp contributeurs Le Bailly 2020 consulte le 24 octobre 2023 Informations lexicographiques et etymologiques de Scarabee dans le Tresor de la langue francaise informatise sur le site du Centre national de ressources textuelles et lexicales Latreille 1802 Histoire naturelle generale et particuliere des Crustaces et des Insectes Tome 3 F Dufart Paris Internet Archive BHL a et b Catalogue of Life Checklist consulte le 23 juin 2023 Stephane Rennesson Emmanuel Grimaud Nicolas Cesard Duel en miniature La delicate mise en scene des combats de scarabees dans le nord de la Thailande Insectes 151 4 3 6 2008 Thiti Wannamontha Jeux Le duel de scarabees une passion thailandaise sur courrierinternational com Courrier international 18 janvier 2012Voir aussiArticles connexes Scarabee du taroLiens externes Sur les autres projets Wikimedia Scarabaeidae sur Wikimedia CommonsScarabaeidae sur Wikispecies en Animal Diversity Web Scarabaeidae consulte le 23 novembre 2023 en Catalogue of Life Scarabaeidae Latreille 1802 consulte le 23 novembre 2023 fr en EOL Scarabaeidae consulte le 23 novembre 2023 en Fauna Europaea Scarabaeidae consulte le 18 juin 2021 fr en GBIF Scarabaeidae consulte le 23 novembre 2023 fr INPN Scarabaeidae Latreille 1802 TAXREF consulte le 23 novembre 2023 en IRMNG Scarabaeidae Latreille 1802 consulte le 23 novembre 2023 fr en ITIS Scarabaeidae Latreille 1802 consulte le 23 novembre 2023 en NCBI Scarabaeidae taxons inclus consulte le 23 novembre 2023 en OEPP Scarabaeidae consulte le 23 novembre 2023 en Paleobiology Database Scarabaeidae consulte le 23 novembre 2023 en Taxonomicon Scarabaeidae Latreille 1802 consulte le 23 novembre 2023 en Tree of Life Web Project Scarabaeidae consulte le 18 juin 2021 en UICN taxon Scarabaeidae consulte le 23 novembre 2023 en WoRMS Scarabaeidae Latreille 1802 liste especes consulte le 23 novembre 2023 Portail de l entomologie